Junco vulcani VOLCANO JUNCO. Thickets, scrubby openings,
bushy areas, bare open ground. High mts. above timberline of Costa
Rica, on Irazú and Turrialba volcanoes, n Cordillera de Talamanca
and w Panama.
Junco hyemalis DARK-EYED JUNCO. Forest, edge, bogs, open
woodland, brushy areas. From Alaska, c,s Yukon, c int. British Columbia,
nw,c Mackenzie, s Keewatin, n,sc Saskatchewan and n Manitoba e across c,s
Canada to Labrador and Newfoundland and s to n,ec Minnesota, se Wisconsin,
c Michigan, ne Ohio, se New York and se New England and in Appalachians
to n Georgia and nw S. Carolina; from sc,se Alaska, coastal and c British
Columbia, wc,s Alberta and sw Saskatchewan s to n Baja Calif., w Nevada,
n Utah, s Idaho and nw Wyoming; se Montana and w S. Dakota s to ne Wyoming
and nw Nebraska; mts. from s Idaho, n Utah and s Wyoming s to e Calif.,
c Arizona, s New Mexico and w Texas.
This species formerly was divided into Slate-colored, Oregon (including
Pink-sided), White-winged and Gray-headed juncos, but these forms intergrade
freely, sometimes forming wide areas where intergrades occur, except where
suitable nesting habitat is restricted. Rarely hybridizes with Zonotrichia
albicollis -- see above under that species.
Junco insularis GUADALUPE JUNCO. Pine, oak, and cypress
forest, woodland. Guadalupe I., off s Baja Calif. Often included
in J. hyemalis, but differs in morphology and vocalizations.
Junco phaeonotus YELLOW-EYED JUNCO. Open, coniferous forest,
pine-oak, scrub, fields. Mts. from ne Sonora, s Arizona, sw New Mexico,
Chihuahua, ne Coahuila, Nuevo León and sw Tamaulipas s to Oaxaca
and w Veracruz; mts. of Baja Calif.; mts. of int. Chiapas; mts. of se Chiapas
and w Guatamala. The isolated populations in s Baja California, Chiapas
and Guatemals have been treated as species.
Passerculus sandwichensis SAVANNAH SPARROW. Grasslands,
tundra, meadows, bogs, farmlands, marshes; salt marshes.
Several subspecies groups are morphologically, genetically and perhaps
vocally distinct and could be treated as species, although all but sanctorum
either hybridize to varying degrees or are connected by intermediate populations.
Following from Rising (1993. Birds of N. Amer. 45:2-3).
P. s. sandwichensis. From w,n Alaska, n Yukon, n Mackenzie,
n Keewatin, n Ontario, islands in James Bay, n Quebec, n Labrador and Newfoundland
s to sw,s Alaska, w to Amukta in the Aleutians and Nunivak I., in coastal
regions s to Monterey Bay in wc Calif., in interior to c Calif., s Nevada,
s Utah, ec Arizona, n New Mexico, c Colorado, Nebraska, Missouri, Kentucky,
e Tennessee, w N. Carolina, w Virginia, w Maryland, se Pennsylvania and
n New Jersey and locally in interior highlands of Mexico s to Guerrero
and Puebla; highlands of sw Guatemala.
P. s. princeps. Nova Scotia, Sable I. and adj. mainland.
P. s. beldingi. Salt marshes of coastal s Calif, n to Morro Bay
region and incl. Channel Is. and Baja Calif. Geographically variable;
many subspecies.
P. s. sanctorum. Islas de San Benito off w Baja California.
Passerculus rostratus LARGE-BILLED SPARROW. Salt marshes.
Nw Mexico from ne Baja Calif. s along coast of Sonora to n Sinaloa and
San Benito I. Winters in coastal salt marshes from s Calif, formerly
incl. Salton Sea, s around both coasts of Baja Calif to Sonora and n Sinaloa.
Studies of mtDNA variation between P. sandwichensis and P. rostratus
suggest specific status, although birds from s Baja California are morphologically
intermediate between rostratus and the beldingi group of the Savannah Sparrow.
Nearly every population in Baja California has a distinct mtDNA profile,
and so has princeps. Perhaps rostratus should be merged with sandwichensis.
Ammodramus maritimus SEASIDE SPARROW. Salt marshes, esp.
Spartina, rushes, tidal reedbeds; marsh prairie. Atlantic-Gulf coasts,
from s New Hampshire and Massachusetts s to ne Florida and from w Florida
w to se Texas; formerly along coast of ce Florida where the nigrescens
subspecies (Dusky Seaside Sparrow), has recently become extinct; marsh
prairie of s Florida.
Avise and Nelson (1989. Science 243:646-648) found that Atlantic Coast
and Gulf Coast populations differ in mtDNA profiles, but the two groups
are not in contact and behavioral differences are unknown.
Ammodramus nelsoni NORTHERN SHARP-TAILED SPARROW. Marshes,
wet meadows.
A. n. nelsoni. From ec British Columbia, s Mackenzie, n,sc Alberta,
c,s Saskatchewan and c,s Manitoba, s to w,se N. Dakota, se S. Dakota and
nw Minnesota.
A. n. alterus. Coastal marshes around James Bay and sw
Hudson Bay in n Ontario (Baillie, 1960. Ont. Field Biol. 14:14-23).
Occurs in early summer as far north as Churchill, Manitoba and may breed
(Jehl and Smith. 1970. Birds of the Churchill region, Manitoba. Publ. #1,
Manitoba Mus. Man & Nature); J. D. Rising, J. S. Greenlaw (pers. comm.).
A. n. subvirgatus. Se Quebec along s shore of St. Lawrence
River (from Montmagny to Point-au-Père), locally on n shore of lower
St. Lawrence estuary and along Atlantic coast from e Quebec, incl. Magdalen
Is., Prince Edward I. and Nova Scotia s to Popham Marsh and Scarborough
Marsh in sw Maine where there is limited contact with A. caudacutus with
some interbreeding. See A. caudacutus. See Greenlaw (1993.
Auk 110:286-303) for evidence for this arrangement.
Ammodramus caudacutus SOUTHERN SHARP-TAILED SPARROW Coastal
marshes. From sw Maine at Popham Marsh and Scarborough Marsh s to
sc New Jersey. If the race A. c. diversus is recognized it
breeds from s New Jersey s at least to Chincoteague on the Delmarva Peninsula,
e Virginia. Old records from Pea I. and Roanoke I. in N. Carolina
were migrants (J. S. Greenlaw, pers. comm.). See A. nelsoni.
Ammodramus leconteii LE CONTE'S SPARROW. Moist grass and
sedge meadows, shrubby tangles on edges of marshes and bogs. From
ec British Columbia, Alberta, Saskatchewan, c,s Manitoba, nc,s Ontario
and wc Quebec s to nc Montana, n N. Dakota, se S. Dakota, nw,e Minnesota,
ne Wisconsin, ne Illinois and n Michigan.
Ammodramus henslowii HENSLOW'S SPARROW. Open fields of grass
and weeds, meadows, shrubby vegetation, esp. in damp areas. From
e S. Dakota, c Minnesota, c Wisconsin, c Michigan, s Ontario, n New York
and c New England s to c Kansas, ne Oklahoma, sw,c Missouri, s Illinois,
c Kentucky, c W. Virginia, e Virginia, w S. Carolina and w,c N. Carolina;
e Texas.
Ammodramus bairdii BAIRD'S SPARROW. Short-grass prairie,
scattered bushes, matted vegetation. From se Alberta, s Saskatchewan
and s Manitoba s to c,e Montana, s S. Dakota, se N. Dakota and wc Minnesota.
Ammodramus savannarum GRASSHOPPER SPARROW. Prairie, old
fields, grasslands, farmlands, savanna. From e Washington, s int. British
Columbia and s Alberta e across s Canada to sw Quebec and n New England
and s to s Calif., c Nevada, n Utah, c Colorado, ne New Mexico, n,sc Texas,
c portions of Gulf states, c N. Carolina and se Virginia; c pen. Florida;
se Arizona and n Sonora; locally in México, Veracruz, Chiapas, Guatemala
in Péten and Caribbean lowlands, Belize, Honduras in int. highlands
and pine savanna, ne Nicaragua in pine savanna, nw Costa Rica and Panama;
Jamaica, Hispaniola and Puerto Rico; Curaçao and Bonaire is.; in
Andes, 2700-3050 m of w Colombia, in Cauca V., and w Ecuador.
Ammodramus humeralis GRASSLAND SPARROW. Savanna, open bushy
areas, forest edge. Lowlands and foothills to 1750 m of Colombia,
Venezuela and Guianas s through n,c,e,s Brazil, extreme se Peru, n,e,se
Bolivia and Paraguay to Uruguay and n Argentina.
Ammodramus aurifrons YELLOW-BROWED SPARROW. Savanna, riverine
scrub, open fields with scattered bushes, second growth, farmlands, often
near water. Lowlands, e of Andes, to 1100 m of e Colombia and s Venezuela
s through e Ecuador and e Peru to n,e Bolivia and w Amazonian Brazil
Sometimes considered conspecific with A. humeralis, but the overlap in
range precludes this treatment.
Xenospiza baileyi SIERRA MADRE SPARROW. Bunchgrass areas,
usually in pine regions. Highlands of c Mexico in Durango, Jalisco,
Morelos and México.
Spizella arborea AMERICAN TREE SPARROW. Open willow, low shrubbery,
scrub conifers, bogs. From n Alaska, n Yukon, n Mackenzie, Banks
I., c int. Keewatin, ne Quebec and Labrador s to s Alaska and w to Bristol
Bay and Alaska Pen., nw British Columbia, se Yukon, wc,s Mackenzie, n Saskatchewan,
n Manitoba, n Ontario, James Bay and c Quebec. Molecular data suggest
that this species is not closely related to the other Spizella species,
but an alternate genus has not been suggested.
Spizella passerina CHIPPING SPARROW. Open forest, edge,
oak woodland, pine-oak, thickets, shrubs, towns. From ec,se Alaska,
c Yukon, c Mackenzie and n Saskatchewan e across c Canada to sw Newfoundland
and s to n Baja Calif., sw,ec Calif., s Nevada, c,se Arizona, through highlands
to nc Nicaragua and to c,e Texas, Gulf Coast and nw Florida; lowland pine
savanna from Belize to ne Nicaragua.
Spizella pallida CLAY-COLORED SPARROW. Shrubby areas, thickets,
shrubbery in meadows, dry pastures, dry scrub. From wc,s Mackenzie,
e British Columbia, nw,c Saskatchewan, n Manitoba and c Ontario s to e
Washington, c Montana, se Wyoming, e Colorado, w Kansas, s Nebraska, n
Iowa, n Illinois, c,se Michigan, s, ne Ontario, sw Quebec and w New York.
Spizella taverneri TIMBERLINE SPARROW. Montane scrub, dwarf
birch. Locally in mts. from sw Yukon, nw, int. British Columbia and
wc Alberta s to mts. of se British Columbia and sw Alberta. Formerly
considered conspecific with S. breweri, but differs in vocalizations, morphology
and ecology.
Spizella breweri BREWER'S SPARROW. Brushland, especially
sagebrush; alpine in e Calif. From cs British Columbia, s Alberta,
sw Saskatchewan and sw N. Dakota s, generally e of Cascades and coast ranges,
to e,s Calif., s Nevada, c Arizona, nw New Mexico, c Colorado, sw Kansas,
nw Nebraska and sw S. Dakota.
Spizella pusilla FIELD SPARROW. Old fields, brushy hillsides,
pastures, thorn scrub, forest edge, fencerows. From nw,se Montana,
s Manitoba, n N. Dakota, c Minnesota, nc Wisconsin, nc Michigan, s Ontario,
sw Quebec, New England and s New Brunswick s, e of Rockies, to e Colorado,
w Kansas, w Oklahoma, c,s Texas, Gulf Coast, n Florida and s Georgia.
Spizella wortheni WORTHEN'S SPARROW. Arid brush, thorn
scrub. Ne Mexico in w Zacatecas and sw Tamaulipas. Recorded
elsewhere in Mexico, possibly migrants. Sometimes considered conspecific
with S. pusilla.
Spizella atrogularis BLACK-CHINNED SPARROW. Chaparral,
sagebrush, arid scrub, brushy hillsides. From cs Oregon, c Calif.,
s Nevada, sw Utah, c Arizona, c New Mexico, w Texas, c Nuevo León
and sw Tamaulipas s to n Baja Calif., sw Calif., se Arizona and in Mexican
highlands to Guerrero, Puebla and nw Oaxaca.
Pooecetes gramineus VESPER SPARROW. Plains, prairie, dry
shrublands, savanna, weedy pastures, fields, sagebrush, arid scrub, open
woodland. From ec,s British Columbia, s Mackenzie, n Alberta and
c Saskatchewan e across s Canada to New Brunswick and Nova Scotia and s
to w Oregon, e,s Calif., c Nevada, sw Utah, n,ec Arizona, c New Mexico
and c U.S. from Colorado e to Tennessee and N. Carolina.
Chondestes grammacus LARK SPARROW. Open situations with
scattered trees and bushes, prairie, forest edge, farmlands orchards, savanna.
W Oregon, e Washington, s int. British Columbia, se Alberta, s Saskatchewan,
s Manitoba, nw,c Minnesota, nw Wisconsin, s Michigan, s Ontario and c Pennsylvania
s to s Calif., mostly w of Sierra Nevada, c Nevada, s Arizona, n Mexico
from ne Sonora to n Tamaulipas, s,e Texas, Gulf states e to Alabama, sc
N. Carolina and w Virginia; very local e of Mississippi V.
Amphispiza bilineata BLACK-THROATED SPARROW. Desert scrub,
thorn brush, mesquite, juniper. From e Washington, sc,se Oregon,
sw Idaho, w,s Colorado, nw Oklahoma and nc Texas s, mostly e of Calif.
deserts, to s Baja Calif, incl. many is., n Jalisco, Guanajuato, Querétaro,
Hidalgo, Tamaulipas and c,s Texas. Differences in allozymes and mtDNA
between small samples of eastern and western subspecies have been reported,
but their significance is unclear -- they may be separate species.
Amphispiza belli SAGE SPARROW. Sagebrush, saltbush brushland,
chaparral.
The nevadensis and belli subspecies groups differ in morphology, ecology
and genetics and generally behave as reproductively isolated species in
areas where both are found (Johnson and Marten 1992. Condor 94:1-19) and
Johnson and Cicero (1991. Proc. 20th. Intl. Orn. Congr.:600-610).
Review by Paul DeBenedictis (Birding 1995. 27:134-137).
A. b. nevadensis. SAGEBRUSH SPARROW. From c int.
Washington, e Oregon, s Idaho, sw Wyoming and nw Colorado s to s Nevada,
sw Utah, ne Arizona and nw New Mexico.
A. b. belli. BELL'S SPARROW. Chaparral. W,sw
Calif., incl. San Clemente I., and n,c Baja Calif. (absent from nw Calif.).
The race canescens is not known to intergrade with belli and differs in
nesting habitat and phenology, but has similar allozymes and mtDNA.
A. b. canescens. SALTBUSH SPARROW. (Atriplex) semi-desert
and desert. S San Joaquin Valley of California, n Mohave Desert and
s Owen's Valley in sc Calif. Occurs within ca. 50 miles of nevadensis
in the Owens Valley without evidence of intergradation between them.
They may be separate species.
Aimophila quinquestriata FIVE-STRIPED SPARROW. Dense, bushy
vegetation, grassy hillsides, mesquite, riparian thickets. From extreme
se Arizona s through e Sonora and w Chihuahua to c Sinaloa, w Durango and
n Jalisco.
The transfer of quinquestriata from Aimophila to Amphispiza by the A.O.U.
Checklist Committee in 1983 lacked a strong foundation; most recent workers
suggest it is better retained in Aimophila.
Aimophila mystacalis BRIDLED SPARROW. Arid scrub, thorn
forest, cactus. Highlands of c Mexico in México, s Puebla,
wc Veracruz to Oaxaca, w of Isthmus of Tehuantepec.
Aimophila humeralis BLACK-CHESTED SPARROW. Arid scrub.
W Mexico from s Jalisco s through Colima, Michoacán, Guerrero, Morelos
and s Puebla to extreme w Oaxaca.
Aimophila ruficauda STRIPE-HEADED SPARROW. Arid scrub,
brushy savanna, thickets bordering on fields. Pacific slope of Mexico
from s Durango and Nayarit s to nw Costa Rica and e to int. in Morelos,
s Puebla and c Guatemala.
Aimophila sumichrasti CINNAMON-TAILED SPARROW. Arid scrub.
Pacific slope of sw Mexico in Oaxaca and extreme sw Chiapas.
Aimophila strigiceps STRIPE-CAPPED SPARROW. Grassland.
Lowlands to 1000 m of se Bolivia, n Argentina and sw Paraguay.
Aimophila stolzmanni TUMBES SPARROW. Arid scrub and woodland.
Pacific lowlands to 1400 m of sw Ecuador and nw Peru, from Tumbes s to
sw Cajamarca and n La Libertad.
Aimophila aestivalis BACHMAN'S SPARROW. Open pine woodland,
brushy hillsides, fields with thickets and brambles. From sc Missouri,
c,ne Illinois, c Indiana, c Ohio, sw Pennsylvania, and c Maryland s to
e Oklahoma, e Texas, Gulf coast and sc Florida; presently rare, absent
or decreasing in n half of breeding range.
Aimophila botterii BOTTERI'S SPARROW. Grassland, savanna,
scattered shrubs, coastal prairie. From se Arizona, sw New Mexico,
n Mexico, w to e Sonora, and extreme s Texas s to Chiapas and Tabasco;
locally in Yucatán, Guatemala, Belize, e Honduras, nw,ne Nicaragua
and nw Costa Rica.
Aimophila cassinii CASSIN'S SPARROW. Open grassland and
short-grass plains with scattered bushes, sagebrush, mesquite, yucca.
From se Arizona, New Mexico, c,ne Colorado, s Wyoming, sw Nebraska, wc
Kansas and w Oklahoma s to n Chihuahua, s Coahuila, n Tamaulipas and c,s
Texas.
Aimophila carpalis RUFOUS-WINGED SPARROW. Open flat grassy
areas with scattered thorn bush, mixed bunchgrass, mesquite or cholla cactus.
Sc Arizona, c,se Sonora and n,c Sinaloa.
Aimophila ruficeps RUFOUS-CROWNED SPARROW. Arid rocky and
hilly areas with brush, scattered shrubs, grassy patches, arid scrub, pine-oak,
coastal sage scrub, pinyon-juniper. From c Calif., incl. Channel
Is., sw Utah, nw,c Arizona, c,ne New Mexico, se Colorado, nw,c Oklahoma,
wc Arkansas and nc Texas s to s Baja Calif. and s to Oaxaca, s Puebla,
wc Veracruz and s Tamaulipas.
Aimophila notosticta OAXACA SPARROW. Arid scrub, brushy
hillsides, oak scrub. Sc Mexico in nw,c Oaxaca and probably sw Puebla.
Aimophila rufescens RUSTY SPARROW. Brush areas, scrub,
pine-oak, oak woodland, forest edge, second growth. Mexico from nc,e
Sonora, w Chihuahua, Sinaloa, nw Durango, Nayarit, Jalisco, Guanajuato,
e SanLuis Potosí and s Tamaulipas s, exc. Yucatán Pen., to
nc Nicaragua and nw Costa Rica.
Torreornis inexpectata ZAPATA SPARROW Dense brush and sawgrass
in swampy regions, coastal scrub. Locally in sw,se Cuba in Zapata
swamp; e Oriente Prov; Cayo Coco, off n Camagüey Prov.
Oriturus superciliosus STRIPED SPARROW Open grassy and
shrubby areas near humid forest or pine-oak. Highlands of Mexico
from e Sonora, Chihuahua, Durango, w Zacatecas, Aguascalientes and San
Luis Potosí s to c Oaxaca, Puebla and Vera Cruz.
Pipilo: This genus is based on Pipilo erythrophthalmus
(Linnaeus 1758). The interspecific hybrids described below show that
erythrophthalmus, ocai and chlorurus are closely related. However,
the Brown Towhee group (aberti, crissalis, fuscus, albicollis) may not
be congeneric with Pipilo, but may be closer to Melozone (Sibley, pers.
opinion based on field observations of behavior).
Pipilo chlorurus GREEN-TAILED TOWHEE. Thickets, chaparral,
shrublands, riparian scrub. Mts. from sw,c Oregon, se Washington,
s Idaho, sw Montana, nw,se Wyoming and nc Colorado s to s Calif, primarily
int., s Nevada, c Arizona, s New Mexico and w Texas.
See below under Hybridization.
Pipilo ocai COLLARED TOWHEE. Thickets, shrubby slopes,
humid forest edge, pine-oak or pine-fir woodland. Mts. of c Mexico
from w Jalisco and ne Colima se through nc Michoacán, Guerrero and
e Puebla to wc Veracruz and c Oaxaca; occurs at higher elevations than
erythrophthalmus where both occur.
Hybridization: The extensive and complex pattern of hybridization
between the morphologically distinct species, P. ocai and P. erythrophthalmus,
was studied by Sibley (1950. Univ. Calif. Publ. Zool. 50:109-194) and (1954.
Evolution 8:252-290); Sibley and West (1958. Auk 76:326-338) and Sibley
and Sibley (Auk 81:479-504). Pure ocai is found alone in Guerrero.
In Oaxaca, where elevational overlap occurs between the two species, there
is no hybridization. In eastern Puebla occasional hybridization occurs
on Mt. Orizaba and the incidence of hybridization increases n to w Veracruz.
From Hidalgo s to México and w to Michoacán, there is a hybrid
cline from erythrophthalmus-like birds in the east to ocai-like birds in
the west. The approximate midpoint is near Morelia. Isolated
mountain-top hybrid populations range from nearly pure ocai in Colima to
nearly pure erythrophthalmus in n Jalisco. On Cerro Viejo, a mountain
on the n shore of Lake Chapala, only hybrids occur and similar populations
are present on other isolated mountains in this region. Pure erythrophthalmus
populations are found n of the s Sierra Madre Occidental and in Chiapas
and Guatemala where ocai does not occur.
The Green-tailed Towhee (P. chlorurus) has hybridized with the Rufous-sided
Towhee as indicated by a photograph in Birding, vol. 26:174, 1994.
The Green-tailed is the northern representative of the Collared Towhee
and it is not surprising that hybrids between Rufous-sided and Green-tailed
should occur. The photograph of the hybrid shows the white throat
and rufous cap of the Green-tailed, characters shared with the Collared
Towhee (P. ocai), with other plumage areas like the Rufous-sided.
Pipilo erythrophthalmus RUFOUS-SIDED TOWHEE. Undergrowth
of open woodland, forest edge, second growth, brushy areas, chaparral,
riparian thickets. From s British Columbia e across sw Canada to
s Saskatchewan and s to s Calif., incl. Channel Is., nw,s Baja Calif, s
Nevada, wc,s Arizona through Mexican highlands to Chiapas and c Guatemala
and e to c Dakotas, nc,w Nebraska, ne,c Colorado, e New Mexico and extreme
w Texas; from s Manitoba e across s Canada to sw Quebec, n New York and
c New England and s to ne Texas, ne,sc Louisiana, Gulf coast, w to Mississippi
and s Florida; Revillagigedo Is. on Socorro I.; highlands of c Mexico in
e Michoacán, México, Distrito Federal, Morelos, s Tlaxcala,
w Puebla and n,c Oaxaca. See Hybridization under P. ocai, above.
Eastern plain-winged (erythrophthalmus) and western spotted-winged (maculatus)
races formerly were treated as species. They differ vocally and a
preliminary report suggests that they have distinct mtDNA profiles, but
they hybridize in suitable habitat across the Great Plains producing a
wide variety of plumage patterns. Eastern populations often have
small white spots on the scapulars, even on Long Island, New York.
Pipilo aberti ABERT'S TOWHEE. Desert scrub, esp. near water,
riparian thickets. From se Calif., se Nevada, sw Utah, c,se Arizona
and sw Mexico se to ne Baja Calif. and nw Sonora.
Pipilo crissalis CALIFORNIA TOWHEE. Brushlands, arid scrub,
chaparral, mesquite, riparian thickets, around human habitation.
Pacific lowlands from sw Oregon s through Calif, w of Sierra Nevada and
sw desert region to s Baja Calif.
Pipilo fuscus CANYON TOWHEE. Brushlands, arid scrub, mesquite,
riparian thickets, around human habitation. From sc Arizona, n New
Mexico, se Colorado, nw Oklahoma and w,c Texas s to Sonora, incl. I. Tiburón,
n Sinaloa, and Mexican highlands to nw Oaxaca, wc Veracruz, Puebla and
sw Tamaulipas.
Allozymes, mtDNA profiles and morphometric studies indicate that P.
fuscus is closer to P. albicollis than to P. crissalis, thus recognition
of fuscus and crissalis as species is warranted. Furthermore, mtDNA
studies demonstrated that P. aberti and P. crissalis are sister taxa, with
the relationship of P. fuscus and P. albicollis less strongly supported.
Pipilo albicollis WHITE-THROATED TOWHEE. Arid scrub, undergrowth
of pine-oak. Mts. of c Mexico in e Guerrero, s Puebla and Oaxaca,
w of Isthmus of Tehuantepec.
Melozone kieneri RUSTY-CROWNED GROUND-SPARROW. Arid scrub,
brush, thickets, undergrowth of open forest. Mexico from se Sonora
s through Sinaloa, w Durango, Nayarit, Jalisco, Guanajuato, Colima, Michoacán,
México, Morelos and Guerrero to sw Puebla and c Oaxaca.
Sometimes considered conspecific with M. biarcuatum and the combined
taxon then called Melozone kieneri, but the name biarcuatum has priority.
Melozone biarcuatum PREVOST'S GROUND-SPARROW. Brush, scrub,
dense undergrowth, meadows, farmlands. Locally in highlands of s
Mexico in Chiapas s to Guatemala, El Salvador, w Honduras and c Costa Rica.
Melozone leucotis WHITE-EARED GROUND-SPARROW. Dense forest
understory, forest edge, scrub, brush. Highlands of Mexico in se
Chiapas s to Guatemala, El Salvador, nc Nicaragua and c Costa Rica.
Arremon aurantiirostris ORANGE-BILLED SPARROW Humid forest,
edge, dense second growth. Lowlands to 1800 m of Mexico on
the Gulf-Caribbean slope from s Veracruz, n Oaxaca, Tabasco and Chiapas
s to Nicaragua, both slopes of Costa Rica and Panama and from w,c,se Colombia
s, w of Andes to w Ecuador and nw Peru and e of Andes through e Ecuador
to ne Peru.
Arremon schlegeli GOLDEN-WINGED SPARROW. Deciduous forest,
edge, arid woodland, scrub, thickets. Lowlands to 1400 m of nc,ne
Colombia and n Venezuela.
Arremon taciturnus PECTORAL SPARROW. Humid forest, second
growth. Lowlands to 1200 m, e of Andes in e Colombia and w Venezuela;
lowlands to 1500 m of s Venezuela and Guianas s through Amazonian e Brazil,
from R. Negro and R. Madeira, Piauí and Pernambuco s to Mato Grosso
and Rio Grande do Sul, to n,e Bolivia and se Peru.
Arremon abeillei BLACK-CAPPED SPARROW. Arid scrub.
Lowlands to 700 m of sw Ecuador and nw Peru on Pacific slope, from Tumbes
and Piura s to Cajamarca; nc Peru in upper Marañón V. of
e Cajamarca.
Arremon flavirostris SAFFRON-BILLED SPARROW. Dense forest
undergrowth. Lowlands to 1400 m of c,se Bolivia, Paraguay, int. e,s
Brazil and n Argentina.
Arremonops rufivirgatus OLIVE SPARROW. Deciduous forest
undergrowth, thickets, thorn scrub, dense second growth, mesquite, riparian
brush. Pacific slope of Mexico from Sinaloa s to Oaxaca; nw Costa
Rica; Gulf-Caribbean lowlands from s Texas, Coahuila, and Nuevo León
s to se Mexico, incl. Yucatán Pen., int. Chiapas and Guatamala in
Petén and Belize.
Arremonops tocuyensis TOCUYO SPARROW. Arid scrub, open
woodland, bushes. Lowlands to 1100 m of ne Columbia in Guajira Pen.,
and nw Venezuela.
Arremonops chloronotus GREEN-BACKED SPARROW. Undergrowth
of open woodland, humid forest edge, second growth, brush, scrub.
Gulf-Caribbean lowlands of Mexico in Tabasco, n Chiapas, Yucatán
Pen., s to n Guatemala, Belize and n Honduras.
Arremonops conirostris BLACK-STRIPED SPARROW. Scrub, dense
undergrowth, forest edge, open woodland, towns. Lowlands to 1600
m of Caribbean slope of Honduras and Nicaragua, both slopes of Costa
Rica and Panama, incl. Coiba and Pearl is., Colombia, w Ecuador, Venezuela,
s to Orinoco and n Bolívar and extreme n Brazil.
Atlapetes albinucha WHITE-NAPED BRUSH-FINCH. Thickets,
brush, humid forest undergrowth. Gulf slope of mts. of Mexico
in e San Luis Potosí, Veracruz, Puebla, n Oaxaca and n Chiapas.
Atlapetes gutturalis YELLOW-THROATED BRUSH-FINCH. Brushy areas,
humid forest edge, undergrowth of pine-oak, dense second growth,
scrub, thickets. Highlands, 800-2600 m of Mexico, in s Chiapas, to
nc Nicaragua, Costa Rica, w Panama and Colombia.
Sometimes considered conspecific with A. albinucha.
Atlapetes pallidinucha PALE-NAPED BRUSH-FINCH. Humid forest
edge, second growth, scrub. Andes, 2400-3600 m of Colombia, extreme
sw Venezuela, w,e Ecuador and nw Peru.
Atlapetes rufinucha RUFOUS-NAPED BRUSH-FINCH. Humid forest
edge, second growth. Mts., 1100-3700 m of Colombia, nw Venezuela,
w,e Ecuador, Peru, on w slope of Andes s to sw Cajamarca and e slope throughout,
and c Bolivia.
Atlapetes leucopis WHITE-RIMMED BRUSH-FINCH. Humid forest
Local on Andean slopes, 2100-3000 m, of Colombia and e,se Ecuador.
Atlapetes pileatus RUFOUS-CAPPED BRUSH-FINCH. Humid forest
undergrowth, pine-oak, dense brush. Highlands of Mexico from w Chihuahua,
Sinaloa, w Durango, Nayarit, Jalisco, Guanajuato, San Luis Potosí,
c Nuevo León and sw Tamaulipas s to Oaxaca, Puebla and w Veracruz.
Atlapetes melanocephalus SANTA MARTA BRUSH-FINCH. Shrubby,
humid forest edge, scrub. Santa Marta Mts., 600-3200 m, of ne Colombia.
Atlapetes flaviceps OLIVE-HEADED BRUSH-FINCH. Humid forest.
E. slope of c Andes, 1300-2200 m, of c Colombia.
Atlapetes fuscoolivaceus DUSKY-HEADED BRUSH-FINCH. Humid
forest, scrub. Upper Magdalena V., 1600-2400 m of se Colombia.
Atlapetes tricolor TRICOLORED BRUSH-FINCH. Mossy forest.
Andean slopes, 300-2000 m of Colombia on Pacific slope, w Ecuador and c
Peru.
Atlapetes albofrenatus MOUSTACHED BRUSH-FINCH. Humid forest.
Andes, 100-2500 m, of Colombia, E. Andes from Norte de Santander s to Cundinamarca,
and nw Venezuela.
Atlapetes schistaceus SLATY BRUSH-FINCH. Humid forest edge,
undergrowth. Mts., 1900-3800 m of Colombia, sw Venezuela s locally
in Andes of nw,e Ecuador; Andes of c Peru.
Atlapetes seebohmi BAY-CROWNED BRUSH-FINCH. Forest.
Andes, 2100 m of s Ecuador; Pacific slope of Andes, 1200-2500 m of nw Peru.
Often considered conspecific with A. nationi, but they are not known
to intergrade.
Atlapetes nationi RUSTY-BELLIED BRUSH-FINCH. Arid woodland,
scrub. W slope of W. Andes, 2000-4000 m, of Peru.
Atlapetes leucopterus WHITE-WINGED BRUSH-FINCH. Forest,
woodland.
The two subspecies groups may be a separate species.
A. l. leucopterus. W. Andes of Ecuador, from Pichincha
s to Chimborazo; Andes, 1000-2500 m of sw Ecuador in El Oro and Loja, and
nw Peru in Tumbes, c,se Piura, Lambayeque.
A. l. paynteri. Andes, 1700-2200 m of cn Peru in ne Piura
and Cajamarca.
Atlapetes albiceps WHITE-HEADED BRUSH-FINCH. Arid scrub,
woodland. Pacific slope, 400-1300 m, of se Ecuador and nw Peru, s
to sw Cajamarca.
Atlapetes pallidiceps PALE-HEADED BRUSH-FINCH. Arid scrub.
Andes, 1900-2100 m of sw Ecuador.
Atlapetes rufigenis RUFOUS-EARED BRUSH-FINCH. Forest edge,
second growth. Andes, 3000-4000 m, of nw Peru in Cajamarca, Libertad,
Ancash, Huánuco; Andes, 2750-4000 m, of s Peru in Apurimac and Cuzco.
Atlapetes semirufus OCHRE-BREASTED BRUSH-FINCH. Humid forest
edge, second growth, scrub, thickets. Mts., 600-3500 m of Colombia
and Venezuela.
Atlapetes personatus TEPUI BRUSH-FINCH. Humid forest, woodland,
shrubs, bushes. Pantepui, 1000-2500m of s Venezuela and extreme n
Brazil.
Atlapetes fulviceps FULVOUS-HEADED BRUSH-FINCH. Dense humid
forest undergrowth. Andes, 1500-3000 m of c,se Bolivia and nw Argentina.
Atlapetes citrinellus YELLOW-STRIPED BRUSH-FINCH. Dense
scrub. Andes, 700-2000 m, of nw Argentina; w Paraguay.
Atlapetes brunneinucha CHESTNUT-CAPPED BRUSH-FINCH. Humid
forest undergrowth, pine-oak, dense second growth, woodland. Mexico
in s Veracruz; locally in mts., 800-3100 m, of Mexico, from e San Luis
Potosí and n Veracruz s to Guerrero, c Veracruz, Oaxaca, Chiapas,
c Guatemala, n El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia,
n Venezuela, Ecuador and se Peru.
Atlapetes virenticeps GREEN-STRIPED BRUSH-FINCH. Humid forest
undergrowth, pine-oak. Mts. of Mexico from s Sinaloa and w Durango
s through Nayarit, Jalisco, Colima and Michoacán to México,
Distrito Federal, Morelos and w Puebla. Sometimes considered conspecific
with A. torquatus.
Atlapetes atricapillus BLACK-HEADED BRUSH-FINCH. Humid
forest edge, dense second growth, thickets. Locally in highlands,
500-1500 m of Costa Rica, Panama and n Colombia.
Sometimes considered conspecific with torquatus, but not known to intergrade.
Atlapetes torquatus STRIPE-HEADED BRUSH-FINCH. Humid forest
edge, second growth, thickets. Mts., 900-3600 m, of Colombia, locally
in Andes, Sierra de Perijá and Santa Marta Mts., and w Venezuela;
mts., 1000-3000 m of n Venezuela; Andes of w,e Ecuador, Peru, w slope s
to sw Cajamarca and e slope throughout, c,se Bolivia and nw Argentina.
Pezopetes capitalis LARGE-FOOTED FINCH. Brushy undergrowth
of humid forest edge, páramo, bamboo thickets, dense second growth.
Mts. of c Costa Rica to w Panama.
Pselliophorus tibialis YELLOW-THIGHED FINCH. Humid forest
growth, bamboo thickets, brush, dense second growth. Mts. of nc Costa
Rica to w Panama.
Pselliophorus luteoviridis YELLOW-GREEN FINCH. Humid forest
undergrowth, edge, brushy areas, usually near water. Mts. of w Panama.
Sometimes considered conspecific with P. tibialis.
Lysurus crassirostris SOOTY-FACED FINCH. Humid forest undergrowth,
edge. Locally in mts., 800-3100 m of c,e Costa Rica, Panama and extreme
nw Colombia. Sometimes included in L. castaneiceps.
Lysurus castaneiceps OLIVE FINCH. Humid forest undergrowth,
edge. Locally in Andes, 700-2200 m, of Colombia, w slope of W. Andes
from Antioquia to Cauca and e slope of E. Andes in se Nariño, nw,e
Ecuador and n,e Peru.
Gubernatrix and Paroaria often are placed in the Cardinalini,
e.g., Ridgely and Tudor (1989). The distinctions among the cardinalines,
emberizines and thraupines are uncertain.
Gubernatrix cristata YELLOW CARDINAL. Shrubbery. Lowlands
to 500 m of s Brazil, Uruguay and ne Argentina.
Paroaria coronata RED-CRESTED CARDINAL. Wet scrub, shrubbery.
Lowlands to 500 m of n,e,se Bolivia, Paraguay, s Brazil, Uruguay and n
Argentina.
Paroaria dominicana RED-COWLED CARDINAL. Open forest, edge,
second growth. Lowlands to 1200 m of e Brazil.
Paroaria gularis RED-CAPPED CARDINAL. Open forest, edge,
second growth, scrub, esp. near water. Lowlands to 300 m of Venezuela
and Trinidad; e of Andes from e Colombia, s Venezuela and Guianas s to
e Ecuador, e Peru, n,e Bolivia and n, Amazonian Brazil.
Paroaria baeri CRIMSON-FRONTED CARDINAL. Second growth,
open woodland, edge. Lowlands to 400 m of c Brazil. Perhaps
conspecific with P. gularis.
Paroaria capitata YELLOW-BILLED CARDINAL. Shrubbery in
humid areas, forest edge, woodland. Lowlands to 500 m of se Bolivia,
sw Brazil, Paraguay and n Argentina. Intro. Hawaiian Is. on Kona
coast of w Hawaii.
Tribe PARULINI: Wood Warblers.
Gill (1994. J. für Orn. 135:359) summarized the well-known
hybridization between V. pinus and V. chrysoptera in eastern North America.
The situation is relatively recent and involves range expansion, hybridization
and replacement of one species by a close relative. Southern V. pinus
has been expanding northward and hybridizing with northern chrysoptera
where they come into contact. Local populations include phenotypically
"pure" individuals of both species and birds that are mixtures of the characters
of both parental types. The composition of populations changes as
the hybrid zone moves northward. Comparisons of mtDNA's suggest that
pinus mtDNA introgresses rapidly into chrysoptera phenotypes without comparable
reverse introgression of chrysoptera mtDNA into pinus. Field identifications
at the species level in or near the hybrid zone should be considered uncertain.
Vermivora bachmanii BACHMAN'S WARBLER. Deciduous woodland,
swamps, probably restricted to bamboo for breeding. Se U.S. in ne
Arkansas, se Missouri, sc Kentucky, c Alabama and se S. Carolina.
Winters in Cuba and Isle of Pines. Possibly extinct; last confirmed
sighting in 1962, but there are recent sight records from Cuba and Louisiana.
Vermivora pinus BLUE-WINGED WARBLER. Brushy hillsides,
second growth, bogs. Se U.S. from e Nebraska, c Iowa, se Minnesota,
s Wisconsin, s Michigan, n Ohio, s Ontario, c New York and New England
s to nw Arkansas, ec Missouri, c Tennessee, n Alabama, n Georgia, w S.
Carolina, w N. Carolina, n Virginia, c Maryland and Virginia. Hybridizes
with V. chrysoptera; see above.
Vermivora chrysoptera GOLDEN-WINGED WARBLER. Deciduous woodland,
second growth, bogs. From ne N. Dakota, s Manitoba, c Minnesota,
n Wisconsin, n Michigan, s Ontario, sw Quebec, n New York, s Vermont and
e Massachusetts s to se Iowa, n Illinois, n Indiana, s Ohio, se Pennsylvania
and s Connecticut and s in Appalachians to n Georgia and nw S. Carolina.
Hybridizes with V. pinus and being replaced by pinus in the southern part
of its range.
Vermivora peregrina TENNESSEE WARBLER. Coniferous and deciduous
woodland, thickets. From se Alaska, s Yukon, and nw,s Mackenzie e
across c Canada to s Labrador and w Newfoundland and s to s Canada, w to
sc British Columbia, nw Montana, n Minnesota, n Wisconsin, n Minnesota,
ne New York and n New England.
Vermivora celata ORANGE-CROWNED WARBLER. Deciduous and mixed
deciduous-coniferous woodland, chaparral, riparian thickets. From
w,c Alaska, c Yukon, nw,s Mackenzie and n Alberta e across c Canada to
s Labrador and s to s Alaska, Alaska Pen. and Kodiak I., sw,c Calif., incl.
Channel Is., Los Coronados and Todos Santos is., off nw Baja Calif., sc
Nevada, c Utah, se Arizona, s New Mexico, and w Texas and e of Rockies
to s Canada, e to sc Quebec.
Vermivora ruficapilla NASHVILLE WARBLER. Open deciduous
and coniferous woodland, second growth, bogs, brushy pine-oak forest.
From sc British Columbia, s Alberta and nw Montana s to nw,c Calif. and
wc Nevada and from c Saskatchewan e across s Canada to Nova Scotia and
s to n,ec Minnesota, s Wisconsin, ne Illinois, s Michigan, ne Ohio, ne
W. Virginia, w Maryland, se Pennsylvania, n New Jersey, se New York and
s New England.
Vermivora virginiae VIRGINIA'S WARBLER. Montane woodland,
oak, pinyon-juniper, chaparral. Sw U.S. from ec Calif., c Nevada,
se Idaho, s Wyoming and nc Colorado s to sc Calif., s Nevada, c,se Arizona,
s New Mexico and w Texas. Sometimes considered conspecific with V.
ruficapilla, but vocally distinct and they are not known to intergrade.
Vermivora crissalis COLIMA WARBLER. Montane thickets, scrubby
woodland, juniper scrub. Cs U.S. and ne Mexico in w Texas in Chisos
Mts., Coahuila, w,c Nuevo León, ne Zacatecas, n San Luis Potosí
and sw Tamaulipas. Sometimes considered conspecific with V. ruficapilla,
but they are vocally distinct and not known to intergrade.
Vermivora luciae LUCY'S WARBLER. Mesquite thickets, scrubby
woodland. Sw U.S. from se Calif., s Nevada, Utah and sw Colorado
s to n Mexico in ne Baja Calif. and n Sonora, s Arizona, s New Mexico and
w Texas.
Parula americana NORTHERN PARULA. Deciduous forest, woodland,
swamp, generally near water. From s Canada, se Manitoba e to Nova
Scotia, s to sc,s Texas, Gulf coast and s Florida; local and sporadic breeding
in New Mexico and coastal Calif.
Dendroica potomac is now regarded as a hybrid P. americana x Dendroica
dominica.
Parula pitiayumi TROPICAL PARULA. Humid forest, edge, woodland,
scrub. Lowlands and foothills to 2600 m from s Texas, in lower Rio
Grande V., and Mexico s to Panama, incl. I. Coiba, and from Colombia, Venezuela
incl. Margarita I., Trinidad, Tobago and Guianas s, w of Andes to nw Peru
and e of Andes, exc. Amazonia, to e Peru, Bolivia, Paraguay, Uruguay, s
Brazil and n Argentina. Revillagigedo Is., on Socorro I.
Parula superciliosa CRESCENT-CHESTED WARBLER. Humid forest,
pine-oak, deciduous woodland. Highlands of Mexico from Sonora, s
Chihuahua, e Sinaloa, w Durango, Nayarit, Jalisco, San Luis Potosí,
c Nuevo León and w Tamaulipas s to nc Nicaragua.
Parula gutturalis FLAME-THROATED WARBLER. Humid forest,
woodland, scrub. Highlands of c,s Costa Rica and w Panama.
Dendroica petechia YELLOW WARBLER. Scrub, second growth,
riparian forest, thickets, towns, esp. near water; mangroves, scrub, thickets.
See note under D. caerulescens re hybridization.
Three well-defined subspecies groups sometimes treated as species.
D. p. petechia and D. p. erithachorides are mangrove specialists.
D. p. aestiva. From Alaska, n Yukon, nw,s Mackenzie and
n Saskatchewan e across c Canada to s Labrador and Newfoundland s to n
Baja Calif., c Mexico, to n Guerrero, Puebla and se San Luis Potosí,
locally in c,ne Texas, Arkansas, e Louisiana, Mississippi, Alabama, Georgia
and c S. Carolina.
D. p. petechia. Cw,s Florida and Bahamas s to Barbados,
islands off Venezuela and Cozumel Island.
D. p. erithachorides. Coasts from s Baja Calif and s Sonora
s along Pacific coast, incl. Cocos I. off Costa Rica, Escudo, Coiba and
Pearl islands off Panama, and Galapagos Islands, to nw Peru; Gulf-Caribbean-Atlantic
coast from s Tamaulipas, incl. Bay Islands off Honduras, to n Colombia
and nw Venezuela.
Dendroica pensylvanica CHESTNUT-SIDED WARBLER. Open deciduous
woodland, edge, second growth, brushy areas. From ec Alberta e across
s Canada to Nova Scotia and s to e Colorado, nc N. Dakota, e Nebraska,
Iowa, n Illinois, n Indiana, c Ohio, e Pennsylvania, c New Jersey and Delaware
and s in Appalachians to c Georgia and nw S. Carolina.
Dendroica magnolia MAGNOLIA WARBLER. Open coniferous and
mixed coniferous-deciduous woodland, edge, second growth. From ne
British Columbia, wc,s Mackenzie and nw Saskatchewan e across s Canada
to s Newfoundland and s to s Canada,w to sc British Columbia, ne Minnesota,
c Wisconsin, c Michigan, nc,ne Ohio, ne Pennsylvania, nw New Jersey and
Connecticut and in Appalachians to w Virginia.
Dendroica tigrina CAPE MAY WARBLER. Open boreal coniferous
forest, edge, woodland. Se Yukon, ne British Columbia, sw,sc Mackenzie
and n Alberta e to Nova Scotia and s to c Alberta, c Saskatchewan, nw N.
Dakota, ne Minnesota, n Wisconsin, s Ontario, ne New York and n New England.
Dendroica caerulescens BLACK-THROATED BLUE WARBLER. Deciduous
upland woodland, second growth. From w,c,s Ontario e to Nova Scotia
and s to ne Minnesota, n Wisconsin, c Michigan, nc,ne Ohio, ne Pennsylvania,
n New Jersey, s New York and s New England and in Appalachians to ne Georgia
and nw S. Carolina. Hybridization with the Yellow Warbler (Dendroica
petechia) reported by Ducharme and Lamontagne (1992. Québec Oiseaux
4:6).
Dendroica coronata YELLOW-RUMPED WARBLER. Coniferous and
deciduous forest, woodland.
D. c. coronata MYRTLE WARBLER, and D. c. auduboni AUDUBON'S
WARBLER, were formerly viewed as separate species, but they intergrade
along the eastern slope of the Canadian Rockies.
D. c. coronata. From w,c,s Alaska, c Yukon, nw,c Mackenzie
and n Manitoba e across c Canada to nc Labrador and Newfoundland and s
to n British Columbia, c,sw Alberta, c,se Saskatchewan, n Minnesota, n
Wisconsin, c Michigan, e Pennsylvania and Massachusetts and s in Appalachians
to W. Virginia and nw Virginia.
D. c. auduboni. From c British Columbia, s Alberta, sw
Saskatchewan, c,se Montana and w S. Dakota s to n Baja Calif., s Arizona,
w Chihuahua, w New Mexico and w Texas; mts. of Mexico from Durango s to
Chiapas and w Guatemala.
Dendroica nigrescens BLACK-THROATED GREY WARBLER. Woodland,
pinyon-juniper, oak scrub. From sw British Columbia, w Washington,
sw Idaho, n Utah, s Wyoming and nw,c Colorado s in mts. to n Baja Calif.,
c,se Arizona, ne Sonora, se New Mexico and w Texas.
Dendroica townsendi TOWNSEND'S WARBLER. Coniferous forest.
From ec,sc,se Alaska, s Yukon, n British Columbia, sw Alberta and sw Saskatchewan
s to nw,c,se Washington, c,ne Oregon, n Idaho, nw,sc Montana and nw Wyoming.
Hybridizes with D. occidentalis in w Washington.
Dendroica occidentalis HERMIT WARBLER. Coniferous forest.
Mts. of w U.S., mostly in Cascades, coast ranges and Sierra Nevada, from
w Washington s to s Calif. and wc Nevada. See townsendi.
Dendroica virens BLACK-THROATED GREEN WARBLER. Open coniferous
and mixed coniferous-deciduous forest, second growth. From n,c Alberta
e across s Canada to s Labrador and Newfoundland and s to n,ec Minnesota,
c Wisconsin, n Indiana, sc,e Ohio, e Pennsylvania, c New Jersey and s New
York and s in Appalachians to c Alabama, n Georgia and w S. Carolina; coastal
plain from se Virginia to e S. Carolina.
Dendroica chrysoparia GOLDEN-CHEEKED WARBLER. Oak-cedar
woodland, scrub, thickets. C Texas, from Dallas Co. s to Edwards
Plateau.
Dendroica fusca BLACKBURNIAN WARBLER. Coniferous forest,
woodland, second growth. From c Saskatchewan e across s Canada to
Nova Scotia and s to c Minnesota, c Wisconsin, c Michigan, ne Ohio, Pennsylvania,
se New York and Massachusetts and s in Appalachians to nc Georgia and nw
S. Carolina.
Dendroica dominica YELLOW-THROATED WARBLER. Pine forest,
sycamore-bald cypress swamp, riparian woodland. From ne N. Dakota,
se Iowa, s Wisconsin, c Indiana, c Ohio, c Pennsylvania, c New Jersey and
Connecticut s to sc,e Texas, Gulf Coast, c Florida and n Bahama Is.
Dendroica graciae GRACE'S WARBLER. Pine forest, pine-oak,
pine savanna. From s Nevada, s Utah, sw Colorado, n New Mexico and
w Texas s in mts. to Nicaragua; lowland pine savanna from Belize
to ne Nicaragua.
Dendroica adelaidae ADELAIDE'S WARBLER. Lowland thickets,
deciduous woodland, forest. C W. Indies on Puerto Rico and n Lesser
Antilles on Barbuda and St. Lucia.
Dendroica pityophila OLIVE-CAPPED WARBLER Pine barrens.
N W. Indies on Cuba and n Bahama Islands.
Dendroica pinus PINE WARBLER. Pine forest, woodland.
From s Manitoba, w,ec Ontario, sw Quebec, c Maine and New Brunswick s to
e Texas, Gulf coast, s Florida, n Bahama Is. and Hispaniola.
Dendroica kirtlandii KIRTLAND'S WARBLER. Scrubby jack-pine.
C Michigan and probably wc Wisconsin and s Ontario. Numbers greatly
reduced.
Dendroica discolor PRAIRIE WARBLER. Brushy second growth,
scrub, mangroves. From e Nebraska, e Kansas, c Missouri, n Illinois,
c Wisconsin, n Michigan, s Ontario, s Pennsylvania, se New York and s New
England s to e Texas, Gulf coast and s Florida.
Dendroica vitellina VITELLINE WARBLER. Scrubby thickets.
Cayman and Swan is. Sometimes considered conspecific with D. discolor.
Dendroica palmarum PALM WARBLER. Bogs, open coniferous
forest, undergrowth near water. From wc,s Mackenzie, ne British Columbia
and n,c Alberta e across c,s Canada to s Labrador and Newfoundland and
s to ne Minnesota, n Wisconsin, c Michgan, n New York and Maine.
Dendroica castanea BAY-BREASTED WARBLER. Boreal coniferous
forest. From sw Mackenzie, ne British Columbia and n,c Alberta, e
across c,s Canada to Newfoundland and Nova Scotia and s to ne Minnesota,
n Wisconsin, n Michigan, ne New York and c New England, with isolated breeding
in Colorado and W. Virginia.
Dendroica striata BLACKPOLL WARBLER. Boreal, coniferous
forest, woodland, alder thickets. From Alaska e across n,c Canada,
n to tree line, to Labrador, Newfoundland and Nova Scotia and s to sc British
Columbia e to nc Ontario and se to e New York, c New England.
Dendroica cerulea CERULEAN WARBLER. Deciduous forest.
From se S. Dakota, se Nebraska, n Iowa, c,se Minnesota, s Wisconsin,
s Michigan, s Ontario, sw Quebec, w,se New York and c New England s to
nc Texas, c portion of Gulf states and c N. Carolina.
Dendroica plumbea PLUMBEOUS WARBLER. Forest, lowland arid
scrub. C Lesser Antilles on Dominica, Marie Galante, Guadeloupe and
Terre-de-Haut is.
Dendroica pharetra ARROWHEAD WARBLER. Montane forest, undergrowth.
Jamaica.
Dendroica angelae ELFIN-WOODS WARBLER. Montane woodland,
humid elfin forest. Highlands of e Puerto Rico.
Catharopeza bishopi WHISTLING WARBLER. Humid forest undergrowth
Mts. of sc Lesser Antilles on St. Vincent I. Sometimes placed in
Dendroica.
Mniotilta varia BLACK-AND-WHITE WARBLER. Deciduous forest,
woodland. Wc,sw Mackenzie, ne British Columbia and n,c Alberta e
across c,s Canada to Newfoundland and s to e Montana, sw S. Dakota, c Nebraska,
c Kansas, sc,e Texas, c portions of Gulf states, and c S. Carolina; c Arizona.
Setophaga ruticilla AMERICAN REDSTART. Open woodland, deciduous
forest, second growth. From se Alaska, s Yukon, wc,s Mackenzie and
Saskatchewan e across c Canada to s Labrador and Newfoundland and s to
nw Calilf., ec Arizona, e Oklahoma, e Texas, c portions of Gulf states
and s S. Carolina.
Protonotaria citrea PROTHONOTARY WARBLER. Swamps, wet lowland
forest. From ec,se Minnesota, sc Wisconsin, s Michigan, s Ontario,
c New York and n New Jersey s to sc,e Texas, Gulf coast and c Florida.
Helmitheros vermivorus WORM-EATING WARBLER. Deciduous forest
undergrowth. From ne Kansas, se Nebraska, n Missouri, se Iowa, c
Illinois, c Indiana, s,ec Ohio, c Pennsylvania, c,se New York and s New
England s to ne Texas, c portions of Gulf states, nw Florida, nw S. Carolina
and ne N. Carolina.
Limnothlypis swainsonii SWAINSON'S WARBLER. Moist forest
undergrowth, canebrakes, swamps. Lowlands and Appalachian highlands
of se U.S. from ne Oklahoma, s Missouri, s Illinois, sw Indiana, sw,e Kentucky,
s Ohio, w W. Virginia, w,s Virginia and s Delaware s to ec Texas, Gulf
coast and n Florida.
Seiurus aurocapillus OVENBIRD. Deciduous forest, woodland,
undergrowth. Ne British Columbia, s Mackenzie, Alberta and c,s Saskatchewan
e across s Canada to Newfoundland and s to s Montana, c,se Colorado, e
Kansas, se Oklahoma, n Arkansas, n portions of Gulf states, w S. Carolina
and c,ne N. Carolina.
Seiurus noveboracensis NORTHERN WATERTHRUSH. Thickets near
water, swamps, bogs. From Alaska, c Yukon, nw,s Mackenzie and n Saskatchewan
e across c Canada to c Labrador and Newfoundland and s to nw Washington,
n Idaho, w Montana, s Saskatchewan, s Manitoba, n N. Dakota, ne Minnesota,
c Wisconsin, c Michigan, ne Ohio, se W. Virginia, Pennsylvania, New York
and Massachusetts; locally nc N. Carolina.
Seiurus motacilla LOUISIANA WATERTHRUSH. Humid forest and
woodland along streams. From e Nebraska, nc Iowa, ec,se Minnesota,
c Wisconsin, s Michigan, s Ontario, c New York and c New England s to e
Texas, s portions of Gulf states and c S. Carolina.
Oporornis formosus KENTUCKY WARBLER. Humid deciduous forest,
swampy woodland. From se Nebraska, c Iowa, sw Wisconsin, ne Illinois,
c Indiana, nc Ohio, s Pennsylvania, n New Jersey, se New York, and sw Connecticut
s to sc,e Texas, Gulf coast, nw Florida, c Georgia and S. Carolina.
Oporornis agilis CONNECTICUT WARBLER. Spruce and tamarack
bogs, poplar woodland. From ec British Columbia e across c Canada
to wc Quebec and s to s Manitoba, n Minnesota, n Wisconsin, c Michigan
and sc Ontario.
Oporornis philadelphia MOURNING WARBLER. Deciduous woodland
undergrowth, second growth, bogs, marsh edges. From ne,c Alberta
e across s Canada to Newfoundland and s to ne N. Dakota, c Minnesota, c
Wisconsin, ne Illinois, s Michigan, n Ohio, ne Pennsylvania, se New York
and c New England and s in Appalachians to w N. Carolina.
Oporornis tolmiei MACGILLIVRAY'S WARBLER. Coniferous forest
undergrowth, riparian thickets, chaparral. From se Alaska, sw Yukon,
n British Columbia, s Alberta, nw Saskatchewan, e Montana and sw S. Dakota
s in mts. to s Calif., c Arizona, s New Mexico and n Nuevo León.
Sometimes considered conspecific with O. philadelphia; they hybridize
occasionally in a contact zone where both are rare, but some reputed intermediates
are within the range of variation in populations far from the contact zone.
Geothlypis trichas COMMON YELLOWTHROAT. Marshes, thickets
near water, brushy pastures, old fields. From se Alaska, s Yukon,
n British Columbia, n Alberta and c Saskatchewan e across s Canada to Newfoundland
and s to n Baja Calif., n,c Mexico s to Veracruz and possibly Oaxaca, s
Texas, Gulf coast and s Florida; Jalisco, around L. Chapala.
A substantial difference in mtDNA profiles has been reported between
eastern and western North American races, but variation in this species
is poorly understood. This, and some or all of the following species
of Geothlypis, are sometimes considered to be conspecific. Most of
these species are allopatric.
Geothlypis beldingi BELDING'S YELLOWTHROAT. Marshes.
S Baja Calif., n to lat 28° N.
Geothlypis flavovelata ALTAMIRA YELLOWTHROAT. Marshes.
Ne Mexico in s Tamaulipas, e San Luis Potosí and n Veracruz.
Geothlypis rostrata BAHAMA YELLOWTHROAT. Brush, scrub,
thickets. N Bahama Is., s to Cat, Long and Little is.
Geothlypis semiflava OLIVE-CROWNED YELLOWTHROAT Tall grass,
bamboo thickets, low bushes, forest edge, woodland, grassy pastures, usually
near water. Lowlands to 1800 m from ne Honduras s, mostly on Caribbean
slope to w Panama; w Colombia and w Ecuador.
Geothlypis speciosa BLACK-POLLED YELLOWTHROAT. Marshes,
wetlands. Highlands of c Mexico in e Michoacán, s Guanajuato,
México and Distrito Federal.
Geothlypis nelsoni HOODED YELLOWTHROAT. Pine-oak undergrowth,
moist brushy areas. From se Coahuila and c Nuevo León s to
Distrito Federal, Puebla, c Veracruz and w,c Oaxaca.
Geothlypis aequinoctialis MASKED YELLOWTHROAT. Damp meadows,
marshes, savanna.
The subspecies groups are as distinct as are other Geothlypis species
and may be separate species.
G. a. costaricensis. Lowlands of sw Costa Rica and w Panama.
G. a. auricularis. Pacific slope from w Ecuador to c Peru.
G. a. aequinoctialis. Lowlands to 1500 m of n,c,e Colombia,
Venezuela, Trinidad, Guianas and n Brazil; e Peru, Bolivia, c,s Brazil,
Uruguay and n Argentina.
Geothlypis poliocephala GREY-CROWNED YELLOWTHROAT. Grassy areas,
brushy fields, thickets. Lowlands of both slopes of Middle America
from n Sinaloa and Tamaulipas s to w Panama.
Microligea palustris GREEN-TAILED GROUND WARBLER. Dense
thickets. Humid mts. and semi-arid lowlands of Hispaniola, incl.
Beata I.
Teretistris fernandinae YELLOW-HEADED WARBLER. Forest undergrowth,
thickets. W Cuba, incl. Isle of Pines and Cayo Cantiles.
Teretistris fornsi ORIENTE WARBLER. Undergrowth, brushy
areas. Humid mts. and semi-arid coasts of e Cuba, incl. Arch. Camagüey.
Leucopeza semperi SEMPER'S WARBLER. Forest undergrowth.
Mts. of c Lesser Antilles on St. Lucia; possibly extinct.
Wilsonia citrina HOODED WARBLER. Deciduous forest.
From se Nebraska, c,ne Iowa, se Minnesota, c Illinois, s Michigan, s Ontario,
nw Pennsylvania, c, se New York and s New England s to e Texas, Gulf coast
and n Florida.
Wilsonia pusilla WILSON'S WARBLER. Bogs, thickets, brushy
areas near water, often in boreal or montane regions. From Alaska,
n Yukon, nw,e Mackenzie and nw,c Saskatchewan e across c,s Canada to c
Labrador and Newfoundland and s mostly in mts. to s Calif., wc,ne Nevada,
sc Utah, sw Colorado and nc New Mexico and s in e to New York, n New England
and Nova Scotia.
Wilsonia canadensis CANADA WARBLER. Woodland undergrowth,
bogs, shrubbery along streams. From n,c Alberta and c Saskatchewan
e across s Canada to Nova Scotia and s to n Minnesota, s Wisconsin, n Illinois,
c Michigan, c Ohio, ec Pennsylvania, n New Jersey, se New York and s New
England and s in Appalachians to nw Georgia.
Cardellina rubrifrons RED-FACED WARBLER. Fir or pine forest,
pine-oak woodland. Mts. from c Arizona, c New Mexico and w Chihuahua
s to w Durango.
Ergaticus ruber RED WARBLER. Pine forest, pine-oak.
Mts. of Mexico in Chihuahua and Durango; from Jalisco s to Guerrero and
e to Hidalgo, e Puebla, c Veracruz and c Oaxaca.
Ergaticus versicolor PINK-HEADED WARBLER. Humid forest,
pine-oak woodland, second growth. Mts. of s Mexico in c,e Chiapas
and w Guatemala. Sometimes considered conspecific with E. ruber.
Myioborus pictus PAINTED REDSTART. Oak, pine forest, pinyon-juniper
woodland, often near water. Mts. of se Calif., nw. c Arizona, sw
New Mexico, w Texas and c Nuevo León s to nc Nicaragua.
Myioborus miniatus SLATE-THROATED REDSTART. Humid forest,
edge, open woodland, pine-oak, second growth. Mts. of Mexico from
s Sonora, s Chihuahua, Durango, Zacatecas and San Luis Potosí s
to Honduras; mts., 500-3000 m of Costa Rica, Panama, Colombia, Venezuela,
w Guyana and extreme nw Brazil s in Andes of Ecuador and nw,e Peru, s on
w slope to Cajamarca, to n,e,se Bolivia.
Myioborus castaneocapillus TEPUI REDSTART. Humid forest,
edge. Pantepui, 1200-2200 m of s,e Venezuela, w Guyana and extreme
n Brazil. Often considered conspecific with M. brunniceps, but vocalizations
are quite different.
Myioborus pariae YELLOW-FACED REDSTART. Humid forest, edge.
Mts., 800-1200 m of ne Venezuela. May be conspecific with M. castaneocapillus;
vocalizations unknown.
Myioborus brunniceps BROWN-CAPPED REDSTART. Humid forest,
edge. Mts. of c, se Bolivia and nw Argentina.
Myioborus albifacies WHITE-FACED REDSTART. Humid forest,
edge. Pantepui, 900-2250 m, of s Venezuela. M. albifacies and
M. cardonai may be conspecific and both may be races of M. castaneocapillus.
Myioborus cardonai SAFFRON-BREASTED REDSTART. Humid forest,
edge. Mts., 1200-1600 m, of s Venezuela.
Myioborus torquatus COLLARED REDSTART. Humid forest, edge,
second growth, thickets. Mts. of Costa Rica and w Panama.
Myioborus melanocephalus SPECTACLED REDSTART. Humid forest,
edge, scrub, woodland. Andes, 2000-3300 m of sw Colombia, w Ecuador,
and nw Peru, s to Cajamarca; e of Andes in Peru and c Bolivia. May
be conspecific with M. ornatus.
Myioborus ornatus GOLDEN-FRONTED REDSTART. Humid forest,
edge. Andes, 1800-3400 m of w,c Colombia; ne Colombia and sw Venezuela.
Myioborus albifrons WHITE-FRONTED REDSTART. Humid forest,
edge. Andes, 2200-3200 m of w Venezuela. May be conspecific
with M. ornatus.
Myioborus flavivertex YELLOW-CROWNED REDSTART. Humid forest,
edge, woodland. Santa Marta Mts., 1500-3000 m, of n Colombia.
May be conspecific with M. ornatus.
Euthlypis lachrymosa FAN-TAILED WARBLER. Tropical deciduous
forest, riparian woodland, second growth, scrub. Mexico from s Sonora,
e San Luis Potosí and s Tamaulipas s on Gulf slope to Veracruz and
n Oaxaca and on Pacific slope to c Nicaragua. May belong in Basileuterus.
Basileuterus fraseri GREY-AND-GOLD WARBLER. Forest, woodland.
Lowlands to 1900 m of w,c Ecuador and nw Peru.
Basileuterus bivittatus TWO-BANDED WARBLER. Humid forest
undergrowth. Pantepui, 900-1800 m, of s Venezuela, Guyana and extreme n
Brazil; Andes, 700-1800 m, of se Peru, Bolivia and n Argentina.
Basileuterus chrysogaster GOLDEN-BELLIED WARBLER. Humid
forest undergrowth. Andes, 300-1200 m, of sw Colombia, on Pacific
slope n to Valle, w Ecuador and c,s Peru.
Basileuterus signatus PALE-LEGGED WARBLER. Humid forest
edge and undergrowth. Locally in Andes in Colombia, c,se Peru, c,se
Bolivia and nw Argentina. Some races of this species may belong in
B. luteoviridis or vice versa.
Basileuterus luteoviridis CITRINE WARBLER. Humid forest,
second growth, bamboo thickets. Andes of w Colombia; c,e Andes, 1700-3650
m of Colombia and sw Venezuela s through Ecuador to se Peru in Cuzco; Andes,
2300-3400 m, of extreme se Peru and wc Bolivia. The southern race
B. l. euophrys is vocally distinct and may be a separate species (J. Curson.
1994. Cotinga 2:50).
Basileuterus nigrocristatus BLACK-CRESTED WARBLER. Humid
forest in shrubby areas and bamboo thickets. Mts., 1300-3400 m, of
Colombia, and w,n Venezuela s to n Peru.
Basileuterus griseiceps GREY-HEADED WARBLER. Humid forest,
second growth. Coastal mts., 1200-1600 m, of n Venezuela.
Basileuterus basilicus SANTA MARTA WARBLER. Humid forest,
edge, bamboo. Santa Marta Mts., 2100-3000 m, of n Colombia.
Basileuterus cinereicollis GREY-THROATED WARBLER. Humid
forest, edge, second growth. Mts., 800-2100 m, of Colombia and w
Venezuela.
Basileuterus conspicillatus WHITE-LORED WARBLER. Humid
forest, second growth. Santa Marta Mts., 750-2200 m, of n Colombia.
Sometimes regarded as conspecific with B. coronatus or B. cinereicollis.
Basileuterus coronatus RUSSET-CROWNED WARBLER. Humid forest
undergrowth, second growth. Andes, 1400-3100 m, of Colombia and sw
Venezuela s to cw Ecuador, Peru and wc Bolivia; n,sw,c,e Ecuador and nw
Peru.
Basileuterus culicivorus GOLDEN-CROWNED WARBLER. Humid
forest undergrowth, edge, deciduous woodland, second growth. Mexico,
from Nayarit, Jalisco, Nuevo León and Tamaulipas s on both slopes
to w Panama; mts., 300-2100 m of w,n Colombia, locally on Pacific slope
and e to Santa Marta Mts., Cesar and Santander and Sierra de Perijá,
and n Venezuela; lowlands to 2100 m, e of Andes, from e Colombia e across
s Venezuela to Trinidad, Guianas and adj. n Brazil; c,s Brazil, n,e Bolivia
and Paraguay to Uruguay and n Argentina.
Basileuterus trifasciatus THREE-BANDED WARBLER. Humid forest
undergrowth, edge. Andes, 500-2000 m, of sw Ecuador and nw Peru.
Sometimes considered conspecific with B. culicivorus.
Basileuterus hypoleucus WHITE-BELLIED WARBLER. Forest undergrowth.
Lowlands to 1000 m of e Bolivia, sc Brazil and ne Paraguay.
Basileuterus rufifrons RUFOUS-CAPPED WARBLER. Forest edge,
riparian scrub, brushy areas. Mts. of Mexico from n Sonora, w Chihuahua,
San Luis Potosí, c Nuevo León and w Tamaulipas s to c Veracruz,
through c Oaxaca and c Chiapas to c Guatemala; Gulf-Caribbean foothills
of s Veracruz, Tabasco, n Oaxaca, n Chiapas, n Guatemala and Belize; lowlands
to 1900 m from s Guatemala, El Salvador and Honduras s to Panama, incl.
Isla Coiba, nc,ne,c Colombia and nw Venezuela. Sporadic and apparently
unsuccessful nesting attempts in s Arizona.
Basileuterus belli GOLDEN-BROWED WARBLER. Humid forest, dense
pine-oak, brushy areas. Mts. of Mexico from Sinaloa, w Durango, México,
e San Luis Potosí and sw Tamaulipas s to c Honduras.
Basileuterus melanogenys BLACK-CHEEKED WARBLER. Humid forest
undergrowth, open woodland, scrub. Mts. of Costa Rica and w Panama.
Basileuterus ignotus PIRRE WARBLER. Humid forest undergrowth,
edge. Mts., 1200-1400 m of extreme e Panama on Cerro Tacarcuna and
extreme nw Colombia in nw Chocó. Often considered conspecific
with B. melanogenys.
Basileuterus tristriatus THREE-STRIPED WARBLER. Humid forest
undergrowth, second growth, thickets. Mts., 300-2700 m of Costa Rica,
w,e Panama, Colombia in Andes and Sierra de Perijá, and n Venezuela
s, w of Andes, to w Ecuador and e of Andes through e Ecuador and e Peru
to c Bolivia.
Basileuterus leucoblepharus WHITE-BROWED WARBLER. Forest
and woodland undergrowth. Lowlands to 1300 m from Brazil s to Paraguay,
s Brazil, e Uruguay and ne Argentina. Has been considered closely
related to B. griseiceps, but may be closer to B. leucophrys.
Basileuterus leucophrys WHITE-STRIPED WARBLER. Forest undergrowth.
Lowlands to 1000 m of c Brazil.
Basileuterus flaveolus FLAVESCENT WARBLER. Deciduous forest
undergrowth, second growth, bushes. Foothills, 200-1200 m, of ne
Colombia and w,nc Venezuela; s Guyana, c,e Brazil, n,e Bolivia and Paraguay.
Basileuterus fulvicauda BUFF-RUMPED WARBLER. Humid forest
along streams, mangroves. Lowlands to 1000 m on Caribbean slope of
c,e Honduras and Nicaragua, both slopes of Costa Rica and Panama and from
w,c,se Colombia s, w of Andes, to nw Peru and, e of Andes, to e Peru,
n Bolivia and extreme w Amazonian Brazil. Sometimes considered conspecific
with B. rivularis.
Basileuterus rivularis NEOTROPICAL RIVER WARBLER. Humid
forest undergrowth near streams, Locally in lowlands to 1400 m, e
of Andes, from e,s Venezuela and Guianas s through e,s Brazil and n,e,se
Bolivia to Paraguay and ne Argentina.
Zeledonia coronata WRENTHRUSH. Dense thickets, bamboo, brushy
areas in humid forest. Mts. of Costa Rica and w Panama. Originally
described as a thrush in 1889 and assigned to the Turdidae or Zeledoniidae
until 1968 when electrophoretic comparisons of egg-white proteins indicated
its true relationships (Sibley, C. 1968. Postilla No. 125, Peabody Mus.
Nat. Hist., Yale Univ.). Zeledonia is closely related to Basileuterus.
Icteria virens YELLOW-BREASTED CHAT. Second growth, riparian
forest undergrowth, thickets, scrub, woodland undergrowth. From s
British Columbia e across s Canada to s Ontario, c New York and c New England
and s to Baja Calif., c Mexico, to Jalisco, México and s Tamaulipas,
Gulf coast and nc Florida.
Granatellus venustus RED-BREASTED CHAT Deciduous forest,
second growth, arid scrub. Pacific lowlands of w Mexico from n Sinaloa
s to Chiapas; Tres Marías Is.
Granatellus sallaei GREY-THROATED CHAT Deciduous forest,
second growth, brushy areas. Lowlands of Mexico from s Veracruz,
Tabasco, n Oaxaca, n Chiapas and Yucatán Pen. s to n Guatemala and
Belize.
Granatellus pelzelni ROSE-BREASTED CHAT. Humid forest,
second growth, esp. near water. Lowlands to 900 m of s Venezuela,
Guyana and Surinam s through Amazonian Brazil to n Bolivia.
Xenoligea montana WHITE-WINGED WARBLER. Forest undergrowth,
thickets. Mts. of Hispaniola.
This species may be a tanager.
Tribe THRAUPINI: Tanagers.
Bananaquit, Conebills, Bush-Tanagers, Hemispingus, Palm-Tanagers, Shrike-Tanagers,
Ant-Tanagers, Mountain-Tanagers, Euphonias, Chlorophonias, Honeycreepers,
Dacnis, Swallow-Tanager, Plushcap, Finch-Tanagers, Sierra-Finches, Diuca-Finches,
Warbling-Finches, Yellow-Finches, Grass-Finches, Grassquits, Seedeaters,
Seed-Finches, Flower-piercers, Orangequit, Ground-Finches, etc.
Like the New World suboscines (Tyranni), the tanagers apparently
evolved in South America during its long isolation from other continents.
The finch-billed tanagers were placed in the Emberizidae or Fringillidae
in the past, but DNA hybridization comparisons showed that they are thraupines
that adapted to the ecological niches in the Neotropics that are occupied
by emberizine and fringilline finches elsewhere. The nectar-feeding
tanagers, including Coereba, Conirostrum, Diglossa, Cyanerpes, Chlorophanes,
Iridophanes, Dacnis, Hemidacnis, Xenodacnis, Ateleodacnis, Oreomanes and
Euneornis were often placed in the family Coerebidae. In other treatments
some of these genera were placed with the Wood Warblers (Parulini) and
other arrangements have been used. The Swallow-Tanager was usually
placed in the monotypic Tersinidae or Tersininae and the Plushcap in the
monotypic Catamblyrhynchidae. The Thraupini herein defined includes
what seem to be the results of an extensive adaptive radiation in South
America that produced ecotypes that resemble the true finches and other
groups. The problem is complicated by the fact that the tanagers
are closely related to the emberizines, parulines, cardinalines and icterines
-- in this classification all are placed in the Emberizinae, but additional
molecular data may modify this classification. Morphological differences
are mainly those associated with feeding, such as bills and tongues, which
are especially subject to convergent evolution. Details of thraupine
phylogeny remain to be clarified. See Bledsoe (1988. Auk 105:504-515)
and Sibley and Ahlquist (1990) for the molecular evidence.
Coereba flaveola BANANAQUIT. Scrub, woodland, second growth,
towns, mangroves, forest, from humid to xeric habitats. Lowlands
in Bahama Is., from Grand Bahama and Little Abaco s to Great Inagua and
Turk Is.; lowlands to 2000 m of W. Indies (but not Cuba and Swan Is.) and
Mexico from c Veracruz, n Oaxaca, Chiapas and is. off Yucatán Pen.,
incl. Holbox, Cancun, Cozumel is. and Cayo Culebra, s in Caribbean lowlands
to Nicaragua, both slopes of Costa Rica and Panama, incl. Coiba and Pearl
is., Venezuela, Trinidad, Tobago and Guianas s, w of Andes, to nw Peru
and, e of Andes, to e Peru, n,e Bolivia, Paraguay, ne Argentina and Brazil.
Seutin, et al. (1994. Evolution 48:1041-1061) analyzed mtDNA restriction-site
variation in Bananaquit population samples from 12 Caribbean islands and
5 localities in Central and n South America. The Jamaican birds are
the most divergent; C. American, n S. American and e Antillean (Puerto
Rico to Grenada) are nearly equally divergent among themselves and may
represent a single, recent range expansion. In the e Antilles three
groups were identified: Puerto Rico, n-c Lesser Antilles (U.S. Virgin Is.
to St. Lucia), and Grenada-St. Vincent. The results suggest a series
of range changes with strong geographic variation over small distances.
The mtDNA data were consistent with several subspecific units, but named
subspecies do not define equally differentiated populations.
Conirostrum speciosum CHESTNUT-VENTED CONEBILL. Gallery
forest, edge, open deciduous woodland, second growth. Lowlands to
1000 m of e Colombia, s Venezuela, Guianas and n Brazil; e Peru, Amazonian,s,se
Brazil, Bolivia, Paraguay, n Argentina.
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