Phyllastrephus xavieri XAVIER'S GREENBUL. Forest.
Patchy distribution in lowlands from s Cameroon, C. African Rep., n,ne
Zaire and w Uganda s to Congo Rep., sc,ce Zaire and nw Tanzania.
Like Icterine Greenbul, but larger.
The following Malagasy species of greenbuls are placed in Bernieria
by some authors.
Olson (1989. Riv. Ital. Orn. 59:183-195) considered the Malagasy species
of Phyllastrephus to be babblers (Timaliini), based on skeletal traits.
Phyllastrephus madagascariensis LONG-BILLED GREENBUL. Forest.
Lowlands to 1800 m, of Madagascar, except s.
Phyllastrephus zosterops SPECTACLED GREENBUL. Forest.
Lowlands to 1200 m, more commonly above 800 m, of nw,e Madagascar. Nest
of moss, plant fibers; hemispherical; near ground. Eggs 3-4, rosy-white,
reddish spots.
Phyllastrephus apperti APPERT'S GREENBUL. Dry forest undergrowth.
Sw Madagascar.
Phyllastrephus tenebrosus DUSKY GREENBUL. Forest undergrowth.
Lowlands of ne,ce Madagascar. 14.5 cm.
Phyllastrephus cinereiceps GREY-CROWNED GREENBUL. Rain forest
undergrowth. Hills ca. 800 m of ce Madagascar. Nest bowl-shaped of moss
on fork of a small branch, near ground. Eggs 3.
Bleda: Bristlebills. Attend army ant swarms.
Green above, yellow below, dusky breast band.
Chappuis and Erard (1993. Zeitschr. Zool. Syst. Evolut.-forsch. 31:280-299)
reviewed species limits in Bleda based on morphology and vocal behavior,
and recognized four species (instead of three), as follows.
Bleda syndactyla COMMON BRISTLEBILL. Forest undergrowth.
Sierra Leone, Liberia, Ivory Coast, Ghana, Nigeria, s Cameroon, Equatorial
Guinea, Gabon, Congo, s C. African Rep., n,ne Zaire, extreme se Sudan,
Uganda, w Kenya, and w Tanzania s to n Angola, nw Zambia and cs,ce Zaire.
Bleda eximia GREEN-TAILED BRISTLEBILL. Forest undergrowth.
Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, s Nigeria, Gulf of Guinea
Is., Cameroon, Equatorial Guinea, Gabon, Congo, Cabinda, Zaire, C. Afr.
Rep., s Sudan, Uganda and w Tanzania.
Bleda notata LESSER BRISTLEBILL. Forest undergrowth.
Bioko and s Nigeria to s Sudan and w Uganda. Includes ugandae.
Formerly considered conspecific with eximia, but structurally and vocally
closer to canicapilla.
Bleda canicapilla GREY-HEADED BRISTLEBILL. Forest undergrowth.
Senegambia, s Mali, Guinea-Bissau, se Guinea, Sierra Leone, Liberia, Ivory
Coast, Ghana, Togo, Benin, and s Nigeria.
Nicator: Nicators. Forests and thickets of w,c,e
Africa. Often viewed as shrikes. Based on skeletal traits,
Olson (1989. Riv. Ital. Orn. 59:183-195) argued that this genus is malaconotine,
but behavior, isozymes (Hannote, et al. 1989. Biochem. Evol. Syst. 15:629-634)
and DNA hybridization data indicate that it is pycnonotid.
Nicator chloris YELLOW-SPOTTED NICATOR. Forest, thickets.
Senegambia, s Mali, Guinea-Bissau, Sierra Leone, se Guinea, Liberia, Ivory
Coast, Ghana, Togo, Benin, Nigeria, s Cameroon, C. African Rep., Equatorial
Guinea, Gabon, Congo, Cabinda, n,ne Zaire, extreme s Sudan, Uganda and
w Tanzania, s,se,ce Zaire and possibly nw Zambia.
Nicator gularis EASTERN NICATOR. Forest, thickets.
S Somalia and c,se Kenya s through Tanzania, c,e Zambia, Malawi, Zimbabwe
and Mozambique to extreme e S. Africa in e Transvaal and e Natal.
Like chloris with grayish crown.
Nicator vireo YELLOW-THROATED NICATOR. Forest. S
Cameroon, Equatorial Guinea, Gabon, Congo, s C. African Rep., n, ne Zaire
and extreme w Uganda, s to c Angola and sw,cs,c Zaire. Like chloris,
but smaller; lores, throat yellow.
Criniger: Bearded Bulbuls. Conspicuous white or yellow throats,
the long feathers of which form "beards" in some species.
Criniger barbatus BEARDED BULBUL. Forest undergrowth.
Se Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, s Nigeria and
Cabinda.
Criniger chloronotus GREEN-BACKED BULBUL. Forest undergrowth.
S Cameroon, Equatorial Guinea, Gabon and Congo s to Congo River mouth,
e to n,ne,ce Zaire and w Uganda.
Similar to barbatus with white throat. Nest a shallow cup of twigs,
moss, lined with strands of a thread-like fungus (Marasmius) and decorated
with long, leafy stems of an epiphytic fern, Microgramma owariensis, around
the rim; placed on green upper leaves of a shrub on bits of wood, dead
leaves, on which the fungi grow and attach the nest to the substrate.
Eggs 1-2, variable, gray to brown; unmarked or with dark markings.
Criniger calurus RED-TAILED BULBUL. Forest undergrowth.
Sw Senegambia, s Mali, Guinea-Bissau, Guinea, Sierra Leone, Liberia, Ivory
Coast, Ghana, Togo, Nigeria to s Cameroon,. Fernando Po I., Gabon, Congo,
C. Afr. Rep., n,ne Zaire, extreme s Sudan, Uganda, s to sw,cw,ce Zaire
Criniger olivaceus YELLOW-BEARDED BULBUL. Lowland primary
forest undergrowth. Patchy distribution in s Mali, sw Senegambia, se Guinea,
sw Mali, Sierra Leone, Liberia, Ivory Coast, s Ghana, Nigeria, Cameroon,
Gabon, Congo, Zaire and C. Afr. Rep.
Criniger ndussumensis WHITE-BEARDED BULBUL. Forest undergrowth.
From s Nigeria, s Cameroon, C. African Rep. and n,ne,ce Zaire, s to Gabon
and Congo Rep. Like calurus with whitish lores. Hybridizes
with calurus in ne Zaire, but the two are sympatric in Cameroon and Gabon
without interbreeding. Considered a race of C. olivaceus by Dowsett
and Dowsett-Lemaire (1993, p. 352), based on "indistinguishable" vocalizations.
Molecular evidence may be required to clarify this complex situation.
Alophoixus: Asian Bearded Bulbuls. Often included in Criniger.
16-24 cm. S,se Asia.
Alophoixus finschii FINSCH'S BULBUL. Forest. Lowlands to
750 m in peninsular Thailand, Malaya, Sumatra and Borneo.
Alophoixus flaveolus WHITE-THROATED BULBUL. Forest, scrub.
Himalayas to 1500 m from ne India and Burma to sw China and nw,sw Thailand.
Alophoixus pallidus PUFF-THROATED BULBUL. Open forest.
Lowlands to 1800 m in s China, s,e Burma, nw,ne Thailand, Cambodia (exc.
sw), Laos and Vietnam.
Alophoixus ochraceus OCHRACEOUS BULBUL. Forest. Highlands,
200-1500 m in s Burma (s Tenasserim), sw,peninsular,se Thailand, sw Cambodia,
s Vietnam, Malaya, w Sumatra and n Borneo from Sarawak to Sabah.
Alophoixus bres GREY-CHEEKED BULBUL. Forest. Lowlands
to 1000 m of Malay peninsula; Sumatra, Java, Bali, Borneo, sw Philippines
on Calamian Is., Palawan and Balabac.
Alophoixus phaeocephalus YELLOW-BELLIED BULBUL. Forest,
second growth. Lowlands to 1000 m in the Malay peninsula, Sumatra
and adj. islands, and Borneo, N. Natuna Is.
Alophoixus affinis GOLDEN BULBUL. Forest. Lowlands
to 800 m on small islands off e Sulawesi (Peleng, Sangihe, Togian, Banggai,
and Sula Is.) and Moluccas (Morotai, Halmahera, Bacan, Obi, Buru, Ambon,
Seram).
Setornis criniger HOOK-BILLED BULBUL. Lowlands of e Sumatra, Bangka
and Borneo.
20 cm. Hooked bill; crestless; brown above, darkest on crown, tail,
wings; white supercilium; black eye and malar stripes; white below, gray
flanks; tail white-tipped.
Tricholestes criniger HAIRY-BACKED BULBUL. (Placed in Hypsipetes
by some authors). Forest, second growth. Lowlands to 1000 m
in the Malay peninsula, Tioman I., Sumatra and Batu Is., Musala and Lingga
Arch., and Borneo and N. Natuna islands.
16 cm. Crestless; olive-brown above; rufescent tail; pale eyering; yellowish
cheeks; white throat; breast yellow-gray; belly and crissum yellow.
Long, hairlike feather shafts from nape over the back.
Iole: Often included in Hypsipetes. S, se Asia.
Iole virescens OLIVE BULBUL. Forest, second growth.
Lowlands to 1000 m in e India, Burma and nw,sw Thailand.
Iole propinqua GREY-EYED BULBUL. Second growth, scrub.
Lowlands to 1000 m in s China, e,s Burma, Thailand and Indochina.
Iole olivacea BUFF-VENTED BULBUL. Forest, woods, second
growth. Lowlands to 1000 m in the Malay peninsula, Sumatra, Batu
Is., Bangka, Belitung, Riau Arch., Borneo, Anambas, N. Natuna and Banggi
islands.
Iole indica YELLOW-BROWED BULBUL. Forest, woods, scrub,
towns. Mainly above 900 m in s India in the W. Ghats from s Maharashtra
and Goa s to Kerala; Sri Lanka. Nest unlike those of other bulbuls:
a flimsy hammock of grass and dead leaves, decorated with moss and cobwebs;
slung in a horizontal fork. Eggs 2-3, white or pinkish with reddish
spots, blotches.
Ixos: Often included in Hypsipetes. Japan to Philippines;
Taiwan; se Asia.
Ixos palawanensis SULPHUR-BELLIED BULBUL. Forest, edge,
second growth. Mts. of Palawan, Philippine Is.
Ixos philippinus PHILIPPINE BULBUL. Forest, edge.
Lowlands to 2000 m of the Philippines, except Palawan.
Ixos rufigularis ZAMBOANGA BULBUL. Forest, edge.
W Mindanao and Basilan, Philippine Is. Similar to philippinus, but
larger with a darker crown.
Ixos siquijorensis STREAK-BREASTED BULBUL. Forest, edge,
second growth. Small islands in the c Philippines: Romblon, Tablas,
Siquijor, formerly on Cebu.
Ixos amaurotis BROWN-EARED BULBUL. Forest, edge, woods,
second growth, towns. Lowlands to 1100 m in Japan from s Hokkaido
s, including many small islands; Izu, Ryukyu, Daito, Bonin and Volcano
is., s Taiwan and Lan Ÿu I.; small is. of n Philippines on Babuyan Is.,
Batan, Fuga and Camiguin Norte.
Ixos everetti YELLOWISH BULBUL. Forest. C, s Philippines
on Samar, Leyte, Panaon, Dinagat, Siargao, Camiguin Sur, c,e Mindanao and
the Sulu Arch. Geographically variable.
Ixos malaccensis STREAKED BULBUL. Forest. Lowlands
to 1000 m in the Malay peninsula, Sumatra, Bangka, Riau and Lingga arch.;
Borneo.
Hemixos: Often included in Hypsipetes. India, s China,
se Asia, Sumatra, Borneo.
Hemixos flavala ASHY BULBUL. Forest, second growth.
Foothills and mts., 300-1800 m in n,e India, Burma, sw China, Thailand
(exc. c,se), n,s Laos, s Vietnam in s Annam; Sumatra; Borneo. Geographically
variable.
Hemixos castanotus CHESTNUT BULBUL. Forest, second growth.
Lowlands to 1000 m in s China, Hainan I., n Vietnam in Tonkin.
Hypsipetes: See Hemixos, Ixos, Iole, Tricholestes, which
are often included in Hypsipetes. Species of Hypsipetes occur in
s, se Asia, Sumatra, Java, Madagascar, Seychelles Is., Comoro Is., Mascarene
Is., and Nicobar Is.
Hypsipetes mcclellandii MOUNTAIN BULBUL. Forest, woods,
second growth. Mts., 900-2600 m in n,e India, s China, Burma, nw,pen.,se
Thailand, n,w Laos, Vietnam in nw Tonkin, s Annam and Malaya. 25
cm.
Hypsipetes virescens SUNDA BULBUL. Forest. Highlands,
800-2400 m, of Sumatra and Java. Related to Mountain Bulbul.
Geographically variable.
Hypsipetes madagascariensis group: The relationships of
the six following species (madagascariensis to nicobarensis) are unclear.
They have been considered conspecific. Only madagascariensis and
parvirostris are sympatric in the Comoro Is.
Hypsipetes madagascariensis MADAGASCAR BULBUL. Forest,
second growth, towns. Madagascar, Aldabra, Glorieuses and Comoro
Is.
Hypsipetes crassirostris SEYCHELLES BULBUL. Forest, second
growth, woods. Seychelles Is. on Mahé, Praslin and Félicité
Similar to madagascarensis.
Hypsipetes parvirostris COMOROS BULBUL. Humid forest.
Highlands above 500 m on the Comoro Is. of Grand Comoro and Mohéli.
Similar to madagascarensis.
Hypsipetes borbonicus REUNION BULBUL. Forest, second growth,
woods. Réunion I. in the Mascarene Islands.
H. olivaceus of Mauritius has red eyes, H. borbonicus of Réunion
has white eyes. They have been treated as subspecies of borbonicus,
but are recognized as separate species by Dowsett and Dowsett-Lemaire (1993,
p. 352).
Hypsipetes olivaceus MAURITIUS BULBUL. Forest, second growth,
woods. Mauritius I. in the Mascarene Islands. See note under
H. borbonicus.
Hypsipetes leucocephalus BLACK BULBUL. Forest, second growth,
towns. Lowlands to 1600 m in sw India, Sri Lanka; foothills and mts.,
1000-3050 m of n Afghanistan, n Pakistan, n,e India, Burma, c,s China,
Hainan, Thailand, Cambodia, Laos, n,c Vietnam and Taiwan.
1. Black form: all black with red bill and feet. 2. White-headed
form: Black with white head and neck; crissum gray with white scale-like
markings.
Hypsipetes nicobarensis NICOBAR BULBUL. Forest, towns.
Nicobar Is.
Hypsipetes thompsoni WHITE-HEADED BULBUL. Forest edge,
second growth. Mts., 900-2100 m, in Burma (exc. w) and nw Thailand.
Neolestes torquatus BLACK-COLLARED BULBUL. Savanna.
Lowlands to 1500 m in s Gabon, Congo, sw,c,ce Zaire and s to nw Angola,
se Zaire and nw Zambia. Affinities uncertain, sometimes placed with the
shrikes.
Back green; forehead to nape gray; a glossy black streak from lores
through eye to side of crown, continued as a broad band or collar across
the chest; chin, throat, ear coverts buffy white; breast to belly whitish;
flanks greenish; underwings golden yellow; eye brown. Nest a frail
cup of grass, plant stems in a bush; eggs 2, pinkish white with dark pink
and rufous spots.
Malia grata MALIA. Mt. forests of Sulawesi in the Celebes.
Affinities uncertain; may be a babbler. See below under Garrulicinae.
Family (Incertae sedis) HYPOCOLIIDAE
Hypocolius ampelinus GREY HYPOCOLIUS. Palms, tamarisks, oases,
towns. Breeds in sw Asia in the Tigris-Euphrates valleys of sw Arabia,
Iraq and s,e Iran. Non-breeding range n to s Turkmenia, e to c India,
s to Red Sea, Egypt, s Arabia.
Affinities uncertain; usually assigned to the Bombycillidae (waxwings
and allies), but there is no substantial evidence for this relationship;
it is considered to be of uncertain status and its assignment to the vicinity
of the Pycnonotidae is tentative. No DNA evidence available.
23 cm. Male gray above, paler below; black lores, mask, nuchal
band, tail tip; primaries black with white tips. Female lacks the
black areas on the head and tips of primaries are dusky. Food: insects
and fruits, including dates. Nest a large, deep cup of plant material,
placed in a tree 1.5-5 meters above the ground and well concealed.
Eggs 3-5, white with grayish markings.
Family CISTICOLIDAE African
Warblers. *Cisticola, Scotocerca, Rhopophilus, *Prinia, Phragmacia,
Oreophilais, Heliolais, Malcorus, Drymocichla, Urolais, Spiloptila, *Apalis,
*Hypergerus, *Eminia, *Camaroptera, Calamonastes, Euryptila. Africa,
s Asia; two species in Australia, one in s Europe. Asterisks* = DNA
hybridization data available.
DNA hybridization comparisons among the genera traditionally placed
in the Sylviidae identified several distinct groups of "warblers", one
of which is composed of the genera listed above. Although the DNA
data included only six of these genera (identified by asterisks*), the
others are thought to be closely related to them. The Cisticolidae
probably originated in Africa, where most species occur today, and spread
from Africa to Asia.
Cisticola: Africa, s Europe, Asia, Australia. Small;
tail short or, if long, graduated; many species have a black subterminal
spot on the rectrices; plumage colors usually shades of brown with tints
of grays, buffs and tawny. Many species are so much alike that identification,
even in the hand, is difficult, but songs, habitat and geographic distribution
are useful in the field. Females are smaller than males; juveniles
are browner and rustier than adults and often yellowish below.
Nests are of dry grass and of three types: 1. Ball-shaped, side
entrance; most species. 2. In a pocket formed by "sewing" together
the edges of large, adjacent leaves of a tree or bush; only Singing Cisticola,
Red-faced Cisticola. 3. Retort or "soda bottle" shape - a ball with
a tubular opening facing upward, placed in tall grass near the ground and
concealed by overhanging grass or composed of living grass stems bound
by cobwebs; only the Zitting Cisticola builds such a nest. Most nests
are lined with plant down which is added to throughout incubation.
Eggs usually 2-4, white or blue with reddish spots; variable.
Cisticola erythrops RED-FACED CISTICOLA. Rank grassland,
undergrowth. Senegambia, s Mauritania, s Mali, Guinea, Burkina Faso, Sierra
Leone, Liberia, Ivory Coast, Ghana, Togo, Benin, Nigeria, s Cameroon, Gabon,
Congo, C. African Rep., n,ne Zaire and extreme se Sudan to w,c Ethiopia
Rwanda, Burundi, Uganda, w,c Kenya, Tanzania, Malawi, Mozambique, ce,se
Zaire, extreme ne Namibia (Caprivi), Angola, Zambia, Zimbabwe, Botswana
and ne S. Africa in n,e Transvaal, Swaziland and Natal.
Cisticola lepe LEPE CISTICOLA. Grassland, undergrowth.
W Angola and se Zaire. Often lumped with erythrops, but they are
sympatric without interbreeding in se Zaire. However, Dowsett and
Dowsett-Lemaire (1993, p. 360) believe that the morphological distinctions
between lepe and cantans are "not sufficient,... to rule out the relationship"
between them. They conclude that "For the moment it [=lepe] should
not be acccepted as a good species." Apparently, further investigation
is required.
Cisticola cantans SINGING CISTICOLA. Grassland, undergrowth.
Senegambia, s Mali, Guinea-Bissau, Guinea, Burkina Faso, Sierra Leone,
Liberia, Ivory Coast, Ghana, Togo, Benin, Nigeria, Niger, Cameroon,
C. African Rep., s, Chad, n,e,sc Zaire, c,s Sudan and w,c Ethiopia, Eritrea,
s through Uganda, Kenya, Rwanda, Burundi, Tanzania, ne Zambia and
Malawi to e Zimbabwe and n Mozambique.
Cisticola lateralis WHISTLING CISTICOLA. Moist woods, forest
edge. Senegambia, Mali, Guinea, Sierra Leone, Liberia, Ivory Coast,
Ghana, Togo, Benin, Nigeria, Cameroon, Gabon, Congo, C. Afr. Rep., s Sudan,
Uganda, Burundi, w Kenya, s Zaire, nw Zambia and Angola.
Cisticola anonymus CHATTERING CISTICOLA. Grassy clearings
in humid forest. In s Nigeria, Cameroon, C. Afr. Rep., Equatorial
Guinea, Gabon, Congo, Zaire and nw Angola.
Cisticola woosnami TRILLING CISTICOLA. Dry woods, grass.
Ne Zaire, c,sw Uganda, sw Kenya, Burundi, Rwanda and Tanzania s to se Zaire,
w Zambia and n Malawi.
Cisticola bulliens BUBBLING CISTICOLA. Grassland with bushes
and small trees, farms. Wc Africa in w Zaire and w Angola.
Cisticola discolor BROWN-BACKED CISTICOLA. Heath, scrubby
areas near forest. Mts., 1000-3350 m of se Nigeria and Cameroon.
Dowsett and Dowsett-Lemaire (1993, p. 360) consider discolor and chubbi
to be conspecific, citing "field experience (including tape playback experiments)
...." Surely, the widely allopatric distributions of these two populations
argue against assuming that reactions to vocalizations necessarily prove
that they are conspecific.
Cisticola chubbi CHUBB'S CISTICOLA. Heath, scrubby areas
near forest. Highlands mostly above 1500 m in w,nw Kenya, w Uganda,
Rwanda, Burundi, e Zaire and nw Tanzania.
Cisticola hunteri HUNTER'S CISTICOLA. Heath, forest glades.
Mts. above 1500 m of ne Africa in w Kenya, Uganda and n Tanzania.
Cisticola nigriloris BLACK-LORED CISTICOLA. Heath, scrubby
areas near forest. Mts., 1500-2750 m of se Africa in n Malawi, ne
Zambia and s Tanzania.
Cisticola emini ROCK-LOVING CISTICOLA. Bare rocky areas,
often overgrown with grasses, bushes and small trees. Locally across
w,c Africa in Guinea, Sierra Leone, s Mauritania, sw Mali, Liberia, Ivory
Coast, Burkina Faso, s Niger, Ghana, Togo, Benin, n,e Nigeria, Cameroon,
ne Chad, C. Afr. Rep., s Sudan, ne Zaire, w Uganda, Rwanda, Kenya and Tanzania
(exc. sw), s to e Malawi and n Mozambique; also in w Angola. The
East African List Committee consider emini conspecific with aberrans (D.
A. Turner, pers. comm.). Dowsett and Dowsett-Lemaire (1993, p. 360)
agree, but note that "They differ only slightly in morphology, and are
allopatric." This seems to be an argument for separate species, rather
than for being conspecific. This is another example of the application
of different species concepts.
Cisticola aberrans LAZY CISTICOLA. Rocky areas, often with
scant vegetation. Se,e Africa from n,e,s Zambia, sw Tanzania and
w Malawi, s through Zimbabwe, e Botswana and s Mozambique to e S. Africa
w to Transvaal and e Cape Province.
Cisticola bodessa BORAN CISTICOLA. Juniper woods.
Ne Africa in s Sudan, s Ethiopia, Eritrea and n Kenya.
Cisticola chiniana RATTLING CISTICOLA. Acacia woods, bushes.
C,e,s Africa from Congo, Zaire, s Sudan, Ethiopia and s Somalia, s to Burundi,
Uganda, Kenya, Tanzania, Angola, Zambia, Malawi, Mozambique, Zimbabwe,
c Namibia, Botswana and S. Africa in n,e Cape Prov., Transvaal, Swaziland,
n Orange Free State and Natal.
Cisticola cinereolus ASHY CISTICOLA. Bushes, light woods.
Ne Africa from extreme se Sudan, n Ethiopia and n Somalia s through Kenya
to ne Tanzania.
Cisticola ruficeps RED-PATE CISTICOLA. Savanna, steppe.
Senegambia, s Mali, Burkina Faso, Sierra Leone, Ivory Coast, n Ghana, Togo,
Benin, Nigeria, Niger, Cameroon, s Chad and c,s Sudan, Kenya and n Uganda
in the Nile Valley to nw Ethiopia and Eritrea. Includes mongalla
which occurs in the upper Nile Valley of s Sudan and n Uganda. The
inclusion of mongalla is supported by the East African List Committee (D.
A. Turner, pers. comm.) and by Dowsett and Dowsett-Lemaire (1993, p. 360).
The race mongalla seems to be in contact with other populations of C. ruficeps.
Cisticola dorsti DORST'S CISTICOLA. Grass steppe, old fields.
Wc Africa in nw Nigeria, n Cameroon and s Chad. Vocally and ecologically
distinct from, and sympatric with, C. ruficeps.
Cisticola rufilatus GREY CISTICOLA. Savanna, acacia steppe.
Gabon, Congo, w Angola, se Zaire, n Zambia and w Malawi s to c Namibia,
Botswana, Zimbabwe and n S. Africa in n Cape Prov. and w Transvaal.
Cisticola subruficapillus RED-HEADED CISTICOLA. Bushes,
scrub. Sw Africa in w Namibia and S. Africa e to e Cape Province
and sw Orange Free State. Reported from sw Angola, but record doubtful.
Cisticola lais WAILING CISTICOLA. Bushes, light woodland.
Locally in c,s Africa in highlands of w Angola, and from s Tanzania, Zambia
and Malawi s through e Zimbabwe and Mozambique to e,s S. Africa in Transvaal,
Swaziland, Orange Free State, Natal and e,cs Cape Province.
Cisticola distinctus LYNE'S CISTICOLA. Bushes, light woodland.
Highlands of e Uganda and c Kenya. Dowsett and Dowsett-Lemaire (1993,
p. 359) treat distinctus as a race of C. lais because their vocalizations
are apparently identical, thus "point clearly to their being conspecific".
However, they also describe morphological differences and the two populations
are allopatric. Are vocalizations more indicative of relationship
than morphology and allopatry? Who can say with absolute certainty?
Cisticola restrictus TANA RIVER CISTICOLA. Bushes, light
woodland. Ne Kenya, along lower Tana River.
Cisticola njombe CHURRING CISTICOLA. Bracken. Highlands
of ec Africa in sc, sw Tanzania, extreme ne Zambia and n Malawi.
Cisticola galactotes WINDING CISTICOLA. Short reeds and
grasses in marshes and swamps.
The two subspecies groups differ vocally and ecologically and may be
separate species.
C. g. galactotes. Senegambia, s Mauritania, s Mali, Guinea-Bissau,
Guinea, Burkina Faso, Sierra Leone, Ivory Coast, Ghana, Togo, Benin, Nigeria,
s Niger, Cameroon, Gabon, Congo, Cabinda, C. Afr. Rep., s Chad, c,s Sudan,
w,c Ethiopia, Eritrea, s Somalia, Zaire, Rwanda, Burundi, Uganda, Kenya,
Tanzania, Malawi, n,e,se Angola, ne Namibia, n Botswana, Zimbabwe, Mozambique
and ne S. Africa in ne Transvaal, coastal Natal, extreme e Cape Prov.
C. g. luapala. Ne,c,s Zambia.
Cisticola pipiens CHIRPING CISTICOLA. Tall reeds and grasses
in swamps and marshes. Angola, s Zaire, ne Namibia, Zambia, sw Tanzania,
Burundi, Zimbabwe, and n Botswana.
Cisticola carruthersi CARRUTHER'S CISTICOLA. Tall reeds
and grasses in swamps and marshes. Ec Africa in ce Zaire, Uganda,
w Kenya, Rwanda, Burundi and nw Tanzania.
Cisticola tinniens TINKLING CISTICOLA. Low sedge growth
in swamps and marshes. In c,ne Angola, se,ce Zaire and w,c Kenya,
Zambia, Botswana, Zimbabwe, w Mozambique and S. Africa (exc. wc,nw Cape
Prov.).
Cisticola angolensis ANGOLA CISTICOLA. Grassland, bushes,
shrubs. See C. robustus.
C. a. angolensis. Highlands of c Africa in Cameroon, Congo Rep.,
cw,c,ne Angola and cs Zaire.
C. a. awemba. Ec Africa in se Zaire, n,c Zambia and extreme
sw Tanzania.
Cisticola robustus STOUT CISTICOLA. Grassland, bushes, shrubs.
Highlands in ne,ce Zaire, extreme se Sudan, w,c Ethiopia, Eritrea, Uganda,
w,sc Kenya, extreme ne Tanzania, Rwanda and Burundi. Dowsett and
Dowsett-Lemaire (1993, p. 360) treat angolensis as a race of robustus.
Cisticola aberdare ABERDARE CISTICOLA. Bushy grassland.
Mts. above 2100 m in Aberdare Mts. of c Kenya.
Cisticola natalensis CROAKING CISTICOLA. Grassland and
savanna, usually with bushes and shrubs. Senegambia, s Mauritania,
s Mali, Guinea-Bissau, Burkina Faso, Guinea, Sierra Leone, Liberia, Ivory
Coast, Ghana, Togo, Benin, Nigeria, Cameroon, Gabon, Congo, Zaire, C. Afr.
Rep., s Chad, Sudan, Ethiopia, Eritrea, Somalia, s to Rwanda, Burundi,
Uganda, Kenya, Tanzania, Malawi, Zambia, Angola, Botswana, Zimbabwe, Mozambique
and e S. Africa s to e Cape Province.
Cisticola fulvicapillus PIPING CISTICOLA. Savanna.
C,s Africa from c Angola, sw,se Zaire, Zambia and sw,sc,ce Tanzania, Malawi,
Zimbabwe, s to s Angola, ne Namibia, n,e Botswana, Mozambique and e S.
Africa in Transvaal, Swaziland, Orange Free State, Lesotho, Natal and s,e
Cape Province.
Cisticola angusticauda TABORA CISTICOLA. Savanna.
Extreme se Uganda, extreme sw Kenya, Rwanda, w,c,s Tanzania, se Zaire and
cn Zambia. Overlaps with fulvicapillus in Zambia, se Zaire and s
Tanzania, and hybridization with fulvicapillus occurs only in Zambia in
a 10 mile wide zone. Dowsett and Dowsett-Lemaire (1993, p. 360) argue
for conspecificity, but Irwin (1991. Bull. Brit. Orn. Club 111:228-236)
treats them as separate species.
Cisticola melanura SLENDER-TAILED CISTICOLA. Savanna, bushes.
Locally in c Africa in ne Angola, s Zaire and extreme w Zambia. Previously
placed in Apalis.
Cisticola brachypterus SIFFLING CISTICOLA. Savanna. Senegambia,
s Mali, Guinea-Bissau, Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana,
Togo, Benin, Nigeria, Niger, Cameroon, Gabon, Congo, Zaire, C. Afr. Rep.,
s Chad and s Sudan to w,c Ethiopia, Eritrea, Somalia, s (exc. forested
w,c) to Rwanda, Burundi, Uganda, Kenya, Tanzania, Malawi, s Angola, s Zambia,
e Zimbabwe and c Mozambique.
Cisticola rufus RUFOUS CISTICOLA. Savanna, open grassland.
Senegambia, s Mali, Burkina Faso, Guinea, Ghana, Togo, Nigeria and n Cameroon
to Lake Chad area and w. C. Afr. Rep.
Cisticola troglodytes FOXY CISTICOLA. Savanna. C.
African Rep., s Sudan, w Ethiopia, nw Kenya, ne Zaire and n Uganda.
Cisticola nanus TINY CISTICOLA. Savanna, steppe. From
extreme s Sudan and s,e Ethiopia, s Somalia and c Kenya to n Tanzania.
Cisticola juncidis ZITTING CISTICOLA. Open grassland.
More than one species may be involved.
From w Europe n to w,s France, Belgium and Netherlands, n Mediterranean
region incl. most islands and n Africa from Morocco e to Tunisia an c Egypt
e through s Turkey and Near East to se Iraq and sw Iran. From Senegambia
e through Mali, Ghana, Niger, n Nigeria, C. African Rep., Sudan and Nile
Valley to Ethiopia and s Arabia, s (exc. forests and deserts) to s S. Africa
(recorded in all African countries except Guinea-Bissau, Liberia, Djibouti,
Socotra I. and Gulf of Guinea Is.). Lowlands to 1200 m from India
w to Indus Valley and n to Himalayan foothills, Nepal, c,ce China, Taiwan,
Japan and Izu and Ryukyu islands, s through se Asia (exc. Cochinchina in
s Vietnam), Nicobar Is., and Indonesia (exc. Borneo) to Philippines and
Lesser Sunda Is. e to Tanimbar Is.; cs New Guinea; locally in n Australia
in coastal N. Territory in the Darwin area and Queensland from Cape York
Pen. s to Normanton area and Fitzroy River.
Cisticola haesitatus ISLAND CISTICOLA. Grassland with bushes.
Socotra I. Has been included in C. juncidis, but seems closer to
C. cherinus.
Cisticola cherinus MADAGASCAR CISTICOLA. Grassland.
Lowlands to 2000 m of Madagascar. Reported to occur on Aldabra, but
not confirmed.
Cisticola aridulus DESERT CISTICOLA. Dry grassland.
Senegambia, s Mauritania, s Mali, Burkina Faso, s Niger, Nigeria, s Chad
and c,s Sudan to Ethiopia, Eritrea, Somalia, s (except forested w,c and
deserts) in Kenya, Tanzania, Angola, Zambia, Mozambique, Namibia, Botswana
and Zimbabwe to s S. Africa in Cape Prov.
Cisticola textrix TINK-TINK CISTICOLA. Short, open, dry
grassland. S Angola, Zambia, s Mozambique and S. Africa from Orange
Free State, Transvaal and Swaziland s to s Cape Prov.
Cisticola eximius BLACK-NECKED CISTICOLA. Open grassland.
Senegambia, s Mali, Burkina Faso, Guinea-Bissau, Sierra Leone, Ivory Coast,
Ghana, Togo to c,se Nigeria; from n Zaire and s Sudan e to n plateau of
Ethiopia and Eritrea, s to Uganda and w Kenya.
Cisticola dambo CLOUD-SCRAPING CISTICOLA. Seasonally flooded
grassland. E Angola, s Zaire and nw Zambia.
Cisticola brunnescens PECTORAL-PATCH CISTICOLA. Open, often
moist grassland. Locally in highlands in w Cameroon, Gabon, Congo
and from w Angola, c,s Zaire, Zambia, Ethiopia, Eritrea, Somalia, Kenya
and Tanzania s to n Botswana, ne Zimbabwe, s Mozambique and ne S. Africa
s to e Cape Province.
Cisticola ayresii WING-SNAPPING CISTICOLA. Open grassland.
Locally from Gabon, Congo, e Zaire, s Sudan, Uganda, Rwanda, Burundi, Kenya,
Tanzania, Malawi, s to Angola, Zambia, Zimbabwe, extreme s Mozambique and
S. Africa w to e Cape Prov. and w Orange Free State.
Cisticola exilis GOLDEN-HEADED CISTICOLA. Open grassland.
More than one species may be involved.
Lowlands and foothills to 1500 m in pen.,n,e India, s China, w,ne,s
Burma, Thailand (exc. sw), Cambodia, s Vietnam in s Annam and Taiwan.
From Sumatra, Java, Borneo and the Philippines (exc. Palawan) e to c,e
New Guinea in the lowlands to 1500 m w to Humboldt Bay, Snow Mts. and Oriomo
River, incl. Manam I., Fergusson and Goodenough is. in the D'Entrecasteaux
Arch., Bismarck Arch., islands in Torres Strait, and coastal n,e,se Australia
from nw W. Australia w to Fortescue River, e to n Queensland on the Cape
York Pen., and s, incl. Great Barrier Reef, to s S. Australia to the Murray
River; King I. in Bass Strait.
Incana incana SOCOTRA WARBLER. Bush country, grassy plains.
Lowlands to 1400 m of Socotra Island. Sometimes placed in Cisticola.
Dowsett and Dowsett-Lemaire (1993, p. 360) cite precedents and field evidence
for the recognition of the genus Incana. Forbes-Watson is cited as
considering it closer to Prinia than to Cisticola.
Scotocerca inquieta STREAKED SCRUB-WARBLER. 10 cm.
Arid thickets, semi-desert. N Africa in Morocco, Algeria, s Tunisia,
Mauritania and n Libya; from e Egypt, Near East and Arabia e through e
Iraq, Iran, to s Kazakhstan, Transcaspia and Afghanistan to w Pakistan.
Pale pinkish brown above; crown streaked; whitish supercilium; whitish
below with fine streaking on throat and upper breast; tail long, rounded,
outer rectrices tipped whitish. Nest a large grass ball with side
entrance; eggs 4-5, white or pink with rufous speckles.
Rhopophilus pekinensis WHITE-BROWED CHINESE WARBLER. Dry bushy
hillsides, tamarisk, long grass. W,n China from sw Sinkiang e through
ne Tsinghai, Kansu, Ningsia and Shensi to Hopeh and s Manchuria
15.5 cm. Above sandy gray streaked with black; crown rufescent streaked
with black; supercilium whitish; white below streaked with chestnut on
sides of neck and breast; flanks buffy; crissum brownish; tail dark gray-brown,
long, graduated; two outer rectrices with white outer webs. Affinities
uncertain, may be a babbler.
Prinia: Prinias. S, se Asia; Africa. 12-18
cm. Small and slender; tail long, graduated, often white-tipped;
carried almost erect; above brownish, rufous, gray, some streaked; below
white, buffy to yellow; some spotted or streaked; some with whitish supercilium;
some with streaked throat or crown.
Prinia burnesii RUFOUS-VENTED PRINIA. Long sarkhan grass
(Saccharum) and elephant- grass, near water. Pakistan on the Indus
R. plains.
Burnesii and cinerascens often treated as conspecific, but they differ
in plumage characters and are widely separated geographically. B.
King (pers. comm.) recommends separate species.
Prinia cinerascens SWAMP PRINIA. E India in w Bihar, W.
Bengal, w Assam, n Bangladesh.
Prinia criniger STRIATED PRINIA. Long grass, bushy hillsides,
forest edge, acacia clumps, thorn forest. Lowlands and foothills to 2450
m in Pakistan, n India, Burma (exc. s), s China; Taiwan.
Prinia polychroa BROWN PRINIA. Grassland, scrub.
Lowlands to 1400 m in sw China, c,e Burma, nw,c,ne Thailand, Cambodia,
c,s Laos and s Vietnam in s Annam; Taiwan; Java.
Prinia atrogularis HILL PRINIA. Grassland, dense low scrub,
forest edge. Mts., 600-2450 m, from ne India, s Tibet, Burma (exc.
c) and s China s through nw Thailand, Laos, Vietnam in Tonkin and s Annam,
and Malaya; Sumatra.
Prinia cinereocapilla GREY-CROWNED PRINIA. Pine forest.
Himalayan foothills in n India from Kashmir e to Bhutan; n Cachar Hills
of s Assam.
Prinia buchanani RUFOUS-FRONTED PRINIA. Arid scrub jungle,
coarse grass, bushy semi-desert. Pakistan (Indus R. plains) and c
India from Rajasthan, Punjab and Uttar Pradesh s to Gujarat, c Mahrashtra,
Madhya Pradesh and w Bihar.
Prinia rufescens RUFESCENT PRINIA. Grassland in open forest
and woods, farms, scrubby hillsides. Lowlands to 1800 m in e India
w to e Nepal and Bangladesh; ne Orissa; sw China in w,s Yunnan, and se
Asia (exc. c Thailand).
Prinia hodgsonii GREY-BREASTED PRINIA. Grassland, thickets,
acacia scrub, bamboo, mangroves, thorn forest. Lowlands to 1500 m
in India n to Himalayan foothills and w to Gujarat, Rajasthan, Punjab and
Kashmir; Sri Lanka; sw China and se Asia (exc. Malay Peninsula).
Prinia gracilis GRACEFUL PRINIA. Thickets, tamarisk jungle,
scrub, coarse grass. From Egypt s to c Sudan, Eritrea, Djibouti and
along the Red Sea coast e to n,ne,se Somalia and Arabia; lowlands to 1000
m from s Turkey and Near East through Iraq, se Iran and sw Afghanistan
to n,c Pakistan and n India, s to Gujarat and the Ganges Valley, e to se
Nepal and lower Brahmaputra Valley in w Assam and Bangladesh.
Prinia sylvatica JUNGLE PRINIA. Low jungle with coarse
grass, esp. ravines. Lowlands to 1000 m in extreme ce Pakistan in
Sind and India n to Punjab and Himalayan foothills and e to Bangladesh;
Sri Lanka.
Prinia familiaris BAR-WINGED PRINIA. Scrub, grass.
Lowlands to 1000 m of Sumatra, Java and Bali.
Prinia flaviventris YELLOW-BELLIED PRINIA. Thick scrub,
heavy grass, swamps. Lowlands to 1200 m in the Indus Valley of e
Pakistan and nw India from Punjab s to the Indus delta; from e India w
to Nepal and W. Bengal, s China, Hainan I., and Taiwan, s through se Asia
(exc. sw,se Thailand) to Sumatra, Nias and Java.
Prinia socialis ASHY PRINIA. Gardens, farms, thorn and
deciduous forest. Lowlands and foothills to 1800 m of India n to
Gujarat, Rajasthan, Punjab and s Kashmir, e to w Assam, Meghalaya, Bangladesh
and Manipur; Sri Lanka.
Prinia subflava TAWNY-FLANKED PRINIA. Acacia steppe, savanna.
From Senegambia e through s Mauritania, s Mali, s Chad and c,s Sudan to
w Ethiopia and sw Somalia; s to s Angola, ne Namibia, n,e Botswana and
e S. Africa in Transvaal, Swaziland, e Orange Free State, Natal and e Cape
Province. This species has been recorded in every subSaharan African
country except Djibouti, Equatorial Guinea and the Gulf of Guinea islands.
Prinia somalica PALE PRINIA. Savanna. From se Sudan,
s Ethiopia and Somalia s to n,e Kenya.
Prinia inornata PLAIN PRINIA. Scrub, tall grass, farms.
Lowlands and foothills to 1200 m from n,e Pakistan, India, s Nepal, Sri
Lanka, c,se China, Hainan I., Taiwan; s to se Asia; Java.
Prinia fluviatilis RIVER PRINIA. Tall riparian grass, marshes.
Niger on the Niger River, Senegambia, Cameroon and the Lake Chad area.
Prinia flavicans BLACK-CHESTED PRINIA. Acacia steppe.
Sw,c Angola, n Namibia, Zambia, Botswana and w Zimbabwe s to c S. Africa
from n Cape Prov. e to Orange Free State and Transvaal. Occasional
hybrids with maculosa.
Prinia hypoxantha SAFFRON-BREASTED PRINIA. Forest edge,
wooded gullies, bracken-briar tangles. Drakensberg and other mts.
of se,e South Africa in se Cape Prov., e Orange Free State, Lesotho, Natal,
n,e Transvaal. Limited intergradation with P. maculosa and treated
as a race of maculosa by some authors.
Prinia maculosa KAROO PRINIA. Savanna, karoo, mountain
scrub. Namibia and S. Africa in w,c Cape Prov., w,c Orange Free State
and Basutoland; e Cape Prov. See P. hypoxantha and P. flavicans..
Prinia molleri SÃO TOME PRINIA. Scrub. Island
of São Tomé, Gulf of Guinea.
Prinia robertsi BRIAR WARBLER. Has been placed in Oreolais.
Bracken, briars, tree heath, forest edge. Mts. above 1350 m in e
Zimbabwe and adjacent sw Mozambique.
Sooty brown above; tail brown, unspotted; chin, throat white; chest,
flanks, crissum brownish; belly white. Nest of stripped grass heads,
domed with large side entrance and attached with cobwebs to leaves; eggs
3, bright blue with brown spots.
Prinia leontica SIERRA LEONE PRINIA. Forest undergrowth,
edge, ravines. Mts., 475-1400 m in s Guinea, e Sierra Leone, Liberia
and sw Ivory Coast.
Prinia leucopogon WHITE-CHINNED PRINIA. Bushes and undergrowth
of forest clearings and edge. E Nigeria, Cameroon, Gabon, Congo,
Zaire, C. African Rep., s Sudan, Uganda, Rwanda, Burundi and nw Kenya,
s to n Angola, n Zambia and w Tanzania.
Prinia bairdii BANDED PRINIA. Dense bushes and shrubs in
forest clearings and edge. Lowlands in se Nigeria, Cameroon, sw C.
African Rep., Equatorial Guinea, Gabon, Congo, s to n Angola, and e through
n Zaire to e Zaire, Rwanda, Burundi, Uganda and nw Kenya. Includes
melanops.
Phragmacia substriata NAMAQUA WARBLER. Formerly in the
genus Prinia. Riparian thornbush among karoo scrub. S Namibia
and w S. Africa in w,c Cape Province and sw Orange Free State.
Dull russet-brown above; neck brownish gray; supercilium and underparts
white; flanks, crissum brownish; tail unspotted. Nest a cup of grass,
leaves, etc., placed low in thick cover; eggs 2-4, blue, speckled with
reddish-brown plus gray undermarkings.
Heliolais erythroptera RED-WINGED WARBLER. Savanna with
tangled grass and bushes. S Mali, Senegambia, Guinea, Sierra Leone,
Liberia, Ivory Coast, Burkina Faso, Ghana, Togo, Benin, Nigeria, Cameroon,
C. African Rep. and n Zaire to s Chad, s Sudan and Ethiopia, s in e Africa
through Uganda, w Kenya, Tanzania, e Zambia and Malawi to Mozambique, e
Zimbabwe; possibly ne S. Africa in ne Transvaal.
Crown, nape gray; back gray-brown; rump pinkish and yellow; wings brownish
with chestnut edgings; chin, throat white; breast buffy; belly white; crissum
tawny; tail graduated with white tips and black subterminal spots.
Nest of fine grass, sewn between leaves; eggs 2, green with pinkish undermarkings.
Malcorus pectoralis RUFOUS-EARED WARBLER. Often placed in
the genus Prinia. Dense low scrub. Namibia, cw,s Botswana and
S. Africa in arid regions e to n,c Cape Prov., extreme w Transvaal, nw
Orange Free State.
Crown streaked pale rufous and blackish; lores, supercilium,face and
ear coverts rufous; back streaked black and buff; throat white; black pectoral
collar; belly buffy white; tail graduated. Nest a neat oval of grass
in a low bush; eggs 3-7, pale blue or white.
Drymocichla incana RED-WINGED GREY WARBLER. Swampy woods,
grass. From ec Cameroon e through n C. African Rep. to se Sudan,
ne Zaire and nw Uganda.
Affinities uncertain, may be closest to Camaroptera. Pale gray
above, wings and tail darker; whitish below; tawny primary patch.
Few records.
Urolais epichlora GREEN LONGTAIL. Forest, esp. wooded ravines,
rarely savanna. Highlands in se Nigeria, sw Cameroon and Fernando
Po I. Affinities unknown; possibly a sylviid.
15 cm. Green above; lores yellow; chin, throat whitish-buff; breast
mottled gray; belly white; crissum yellow; tail dark brown, long (7-8 cm),
graduated, rectrices narrow, buff-tipped. Nest, eggs unknown.
Spiloptila clamans CRICKET LONGTAIL or SCALY-FRONTED WARBLER.
Acacia scrub in desert edge. SubSaharan Africa (Sahel) from Mauritania
e through Mali, Burkina Faso, Niger, n Nigeria, Lake Chad region and Sudan
to n Ethiopia and Eritrea. Affinities uncertain; may be a sylviid.
10 cm. Reddish-brown above; rump yellow; pale whitish-buff below;
forehead, wing coverts black and white; tail 4 cm, graduated, gray with
white tips, black subterminal bars. Nest deep, ovate, domed or semi-domed,
of grass; in a small shrub near the ground; eggs 3, white with fine red
speckles.
Spiloptila rufifrons RED-FRONTED WARBLER. Scrubby hillsides
with sparse grass. Chad, Sudan, Ethiopia, Eritrea, Djibouti and Somalia,
s through e Uganda and Kenya to ne Tanzania. Placed in Spiloptila
by the East African List Committee (D. A. Turner, pers. comm.) and by Dowsett
and Dowsett-Lemaire (1993). Has been in Apalis.
Phyllolais pulchella BUFF-BELLIED WARBLER. Acacia savanna,
scrub. N Nigeria, n Cameroon, C. Afr. Rep., Lake Chad area to ne
Zaire, se Sudan, w,c Ethiopia, Eritrea, Rwanda, Uganda, w,c Kenya, n Tanzania.
Pale olive-green above; pale yellow below; tail blackish with white
tips and white outer webs to outer rectrices. Food insects gleaned
from top foliage of small acacias. Nest purse-shaped, woven, mainly
of plant down, hanging from branches. Eggs 2-3, pale greenish-blue
with reddish, gray-brown, or brown spots. Has been placed in the
Sylviidae.
Apalis: Apalis Subsaharan Africa. Relatively long
bills and tails; plumages variable, some with bright colors, black breast
bands; tail graduated, often with white-tipped rectrices. Usually
inhabit woodland undergrowth, some in upper foliage. Food insects.
Nest domed with side entrance or a deep cup or purselike structure, often
of moss. Eggs variable, blue or green with reddish speckles, white,
olive, etc.
The species described as Orthotomus moreaui may be a species of Apalis
and it is assigned to Apalis by the East African List Committee.
See Orthotomus moreaui below under Orthotomus.
Apalis pulchra BLACK-COLLARED APALIS. Humid forest.
Locally in highlands in e Nigeria, Cameroon, sw C. African Rep., se Sudan,
ne,se Zaire, sw Uganda, Rwanda, Burundi and w,c Kenya.
Apalis ruwenzorii COLLARED APALIS. Humid forest.
Mts. of e Zaire, w Uganda and Rwanda.
Apalis thoracica BAR-THROATED APALIS. Forest. Morphologically
variable with isolated populations that may be separate species; intergradation
usually occurs between races where in contact.
A. t. griseiceps. Se Kenya in the Chyulu Hills, n,c Tanzania from
Mt. Kilimanjaro s to Uluguru Mts., s Mozambique along Zimbabwe border and
S. Africa in c,e Transvaal, Swaziland, Natal, e Orange Free State and e
Cape Province.
A. t. fuscigularis. Se Kenya in the Teita Hills.
A. t. murina. In ne,sw Tanzania in the Usambara Mts. and
extreme sw, Malawi, c,e Zambia, Zimbabwe, adj. wc Mozambique (n of griseiceps),
se Botswana and nc S. Africa in w Transvaal.
A. t. flavigularis. Nc Mozambique, just east of the Rift
Valley.
A. t. lynesi. Ne Mozambique on Mt. Namuli.
A. t. thoracica. S S. Africa in sw,sc Cape Province.
Apalis nigriceps BLACK-CAPPED APALIS. Humid forest, second
growth. Lowlands to 1500 m in Sierra Leone, Liberia, Ivory Coast, s Ghana,
s Nigeria, s Cameroon, Fernando Po I., Equatorial Guinea and Gabon to ne,ce
Zaire and Uganda.
Apalis jacksoni BLACK-THROATED APALIS. Forest. Locally,
mostly in highlands to 2450 m in w,s Cameroon, nw Angola, cn,e Zaire,
Uganda, Rwanda, Burundi, extreme se Sudan, w,c Kenya e to Mt. Kenya and
nw Tanzania.
Apalis chariessa WHITE-WINGED APALIS. Forest. Locally
in lowlands of se Kenya on the lower Tana River; up to 1500 m in c,e Tanzania
in the Udzungwa and Uluguru mts., s Malawi and adjacent Mozambique.
Apalis binotata MASKED APALIS. Forest. Locally in
lowlands to 1800 m in Cameroon, Gabon, nw Angola; ne Zaire, w,e Uganda,
nw Tanzania.
Apalis personata BLACK-FACED APALIS. Forest. Locally
in highlands, 1500-2800 m in e Zaire, Uganda, Rwanda and Burundi.
Apalis flavida YELLOW-BREASTED APALIS. Open forest, acacia
steppe, savanna. Senegambia, Sierra Leone, n Ivory Coast, Burkina
Faso, s Ghana, Togo, Benin, Nigeria, Gabon, Congo, Zaire, s C. African
Rep., s Sudan, s Ethiopia, Somalia, Rwanda, Burundi, Uganda, c Kenya, Tanzania,
s (exc. Congo forest region) in Malawi, Zambia, Angola, n Namibia, n Botswana,
n,e Zimbabwe, Mozambique and e S. Africa w to c Cape Province.
Apalis viridiceps BROWN-TAILED APALIS. Dry acacia steppe,
savanna. S Ethiopia, Somalia, n Kenya. The East African List
Committee includes viridiceps in flavida (D. A. Turner, pers. comm.).
Apalis ruddi RUDD'S APALIS. Riparian thickets, bush clumps
in dry woods, humid coastal forest. S Malawi, s Mozambique and e
S. Africa in Swaziland and n Natal.
Apalis sharpii SHARPE'S APALIS. Humid forest. Mts.
in Guinea, Sierra Leone, Liberia, Ivory Coast and Ghana.
Apalis rufogularis BUFF-THROATED APALIS. Forest.
A. r. rufogularis. Mts. in s Nigeria, s Cameroon, Fernando Po
I., Equatorial Guinea, w Gabon and Congo.
A. r. nigrescens. C,e Zaire, s Sudan, Uganda, Rwanda, Burundi,
w Kenya, Tanzania s to nw,ne Angola, nw Zambia and s Zaire.
Apalis argentea KUNGWE APALIS. Forest. Mts. of ce
Zaire; mts., 1200-2100 m in se Zaire and w Tanzania. The East African
List Committee treat argentea as a race of rufogularis (D. A. Turner, pers.
comm.).
Apalis bamendae BAMENDA APALIS. Humid forest. Mts.
of Cameroon; possibly in C. African Rep.
Apalis goslingi GOSLING'S APALIS. Humid forest. Mts.
in s Cameroon, C. African Rep., Gabon, wc,c,se,ne Zaire and ne Angola.
Apalis porphyrolaema CHESTNUT-THROATED APALIS. Humid forest.
Mts. from ce,ne Zaire, Uganda, Rwanda, Burundi and w,c Kenya s to c Tanzania.
Apalis kaboboensis KABOBO APALIS. Humid forest. Mt.
Kabobo, 1660 m, in se Zaire. Known only from the type locality.
Treated as a race of A. porphyrolaema by Dowsett and Dowsett-Lemaire (1990),
but they are allopatric. Either treatment can be defended.
Apalis chapini CHAPIN'S APALIS. Humid forest. These
allopatric forms may be separate species.
A. c. strausae. Mts. in sw Tanzania, Malawi and e Zambia.
A. c. chapini. Uluguru Mts. in ec Tanzania.
Apalis melanocephala BLACK-HEADED APALIS. Humid forest.
Lowlands to 1500 m from c,se Kenya and s Somalia, s through c,e Tanzania
and s Malawi to Zimbabwe and Mozambique.
Apalis chirindensis CHIRINDA APALIS. Humid forest.
Mts. in e Zimbabwe and wc Mozambique; occurs above 1500 m where sympatric
with A. melanocephala.
Apalis moreaui LONG-BILLED APALIS Forest undergrowth in
Tanzania; canopy in Mozambique. Locally in mts. in ne Tanzania and
wc Mozambique. Described as an Orthotomus, but vocalizations suggest this
species is an Apalis and the East African List Committee and Dowsett and
Dowsett-Lemaire (1993) place it in Apalis, "but with some misgivings".
Nest undescribed, which would be helpful in determining its correct affinities.
Apalis cinerea GREY APALIS. Forest. Locally in mts.
in se Nigeria, s Cameroon, Bioko I., Gabon, cw Angola, e Zaire, Uganda,
Rwanda, Burundi, se Sudan, w,nc,c Kenya, Zambia and Malawi.
Apalis alticola BROWN-HEADED APALIS. Forest, riparian woods.
Mts., 1200-2100 m in sw,nw,nc Angola, se Zaire, n Zambia, n Malawi and
n,sw Tanzania.
Apalis karamojae KARAMOJA APALIS. Forest in mts. of n Uganda;
acacia steppe in mts. in ne Tanzania.
Hypergerus atriceps ORIOLE WARBLER. Gallery forest, often
in palms. Senegambia, sw Mali, Burkina Faso, Guinea-Bissau, Guinea
Sierra Leone, Liberia, Ivory Coast, Ghana, Togo, Benin, Nigeria, Niger,
Cameroon , c C. African Rep. and s Chad. Has been assigned to the
Timaliini (babblers), but DNA hybridization evidence indicates it is a
cisticolid.
20 cm. Head, throat, upper breast black, feathers edges white;
yellowish-olive above; bright yellow below; tail long, olive, graduated.
Eminia lepida GREY-CAPPED WARBLER. Papyrus beds, elephant grass,
woodland thickets. Mts. to 2450 m in ne Zaire, extreme se Sudan,
Uganda, Rwanda, Burundi, w,c Kenya and nw, nc Tanzania.
DNA hybridization evidence indicates that Eminia is a cisticolid.
Forehead, crown gray; black band from lores, above eye, around
occiput; bright green above; shoulders rufous; below pale gray; flanks
green; rufous throat patch. Nest of grass, fibers, domed with side
entrance; suspended from or placed in creepers or dense foliage; eggs 2-3,
white with purplish-brown markings, or glossy pale blue.
Camaroptera: Camaropteras. Subsaharan Africa. Plumages green,
olive-green or gray above; white below; superciliaris has a yellow supercilium;
tail short, square-tipped. Feed on insects. Nest of fine grass,
plant down, cobwebs; purse-shaped, between large leaves which are sewn
together; may have a leaf sewn on as a roof; in a dense thicket near the
ground. Eggs 2-3, white, pale blue, greenish blue, with reddish speckles.
Camaroptera brachyura GREEN-BACKED CAMAROPTERA. Humid forest
undergrowth, thickets.
The three races of brachyura have been treated as separate species,
but Dowsett and Dowsett-Lemaire (1993) and the East African List Committee
(D. A. Turner, pers. comm.) consider them to be conspecific. C. brachyura
occurs in every mainland country in subSaharan Africa.
C. b. brachyura. Lowlands from se Kenya, e,s Tanzania,
Zanzibar and Mafia is., and Malawi s through Mozambique and e Zimbabwe
to e S. Africa w to c Cape Province.
C. b. brevicaudata. GREY-BACKED CAMAROPTERA. From
Senegambia, sw Mali and Guinea-Bissau e to c Sudan, Ethiopia and Somalia;
s (exc. nw Angola) to Namibia, Botswana, extreme ne S. Africa in ne Transvaal,
and Mozambique.
C. b. harterti. HARTERT'S CAMAROPTERA. Nw Angola.
Camaroptera superciliaris YELLOW-BROWED CAMAROPTERA. Humid
forest. Lowlands in Guinea, Sierra Leone, Liberia, Ivory Coast, Ghana,
Togo, Nigeria, s Cameroon, Gulf of Guinea Is., Equatorial Guinea, Gabon,
Congo, sw C. African Rep., n,e Zaire and Uganda, and s to nw Angola, sw,cs
Zaire.
Camaroptera chloronota OLIVE-GREEN CAMAROPTERA. Humid forest.
Lowlands in Senegambia, s Mali, Guinea, Sierra Leone, Liberia, Ivory Coast,
Ghana, Togo, Benin, Nigeria, s Cameroon, Gulf of Guinea Is., Equatorial
Guinea, Gabon, Congo, sw C. African Rep., ne,ce Zaire, Rwanda, Uganda,
extreme se Sudan and nw Kenya, s to c Zaire and extreme nw Tanzania.
Calamonastes: Wren-Warblers. Woods, thornbush, mopane,
brachystegia. C, s Africa. Plumage russet, brownish, or gray
above; buffy, white or gray below with black, brownish or dusky bars; tail
short, constantly "fanned". Feed on insects on or near the ground.
Nest of dry grass or fine, silky fibers; oval or round with side entrance;
enclosed in living leaves which are pierced and sewn together with cobweb.
Eggs 2-4, bluish-white with brownish, grayish speckles. Considered
a subgenus of Camaroptera by Dowsett and Dowsett-Lemaire (1993).
Calamonastes simplex GREY WREN-WARBLER. Woodland, esp. mopane
and brachystegia, thorn bush. Extreme se Sudan, Ethiopia, Somalia,
Kenya (exc. extreme sw), ne Uganda and ne Tanzania.
Calamonastes undosus PALE WREN-WARBLER. Woodland.
Se Zaire, n Zambia, n Malawi, Tanzania (exc. ne), Rwanda, Burundi, Mozambique,
Namibia, Botswana, Zimbabwe and ne S. Africa. Includes stierlingi
and cinerea, following Dowsett and Dowsett-Lemaire (1993).
Calamonastes fasciolatus BARRED WREN-WARBLER. Thornbush.
From wc Angola, Namibia, Botswana and sw Zimbabwe to n S. Africa in n Cape
Prov., nw Transvaal and Orange Free State.
Poliolais lopezi WHITE-TAILED WARBLER. Forest undergrowth.
Mts. in wc Africa in se Nigeria, Cameroon Mt., Manenguba and Kupé
mts., and Bioko Prov. on Fernando Po I., Gulf of Guinea.
10 cm. Geographically variable; male grayish/olive or brownish/olive
above; chin, throat gray to olive; below pale olive-green/whitish, flanks
ashy; crissum buff; tail short, center rectrices brown, outer rectrices
white; female rufous head, olive-green back; pale yellowish below. Sometimes
placed in Camaroptera or Orthotomus. Placed here by Dowsett and Dowsett-Lemaire
(1993).
Graueria vittata GRAUER'S WARBLER. Forest. Mts. of
ce Zaire, Congo and Burundi. Affinities uncertain. Placed here
by Dowsett and Dowsett-Lemaire (1993).
Euryptila subcinnamomea KOPJE WARBLER. Desert scrub
around stony hills (kopjes). Nw S. Africa in w,c Cape Province, sw
Botswana and sw Namibia in Karoo Desert.
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