Hirundo semirufa RUFOUS-CHESTED SWALLOW. Open grassland,
savanna. From Senegambia and Sierra Leone e through s Mali, s Niger
and s Chad to cw,s Sudan and w Kenya, coastal Ivory Coast and Ghana, and
s to nc Namibia, n,e Botswana and ne S. Africa in n,e Cape Prov., Transvaal,
Swaziland, Orange Free State and n Natal.
Hirundo senegalensis MOSQUE SWALLOW. Savanna woodland, forest
clearings, towns. From Senegambia e through s Mali, s Niger, s Chad
and c,s sudan to w,c Ethiopia and s Somalia, s incl. Mafia I. off Tanzania,
to n Namibia, n Botswana, Zambia, n,e Zimbabwe, s Mozambique and ne S.
Africa in e Transvaal and n Natal.
Hirundo daurica RED-RUMPED SWALLOW. Open country, towns.
Some of the allopatric populations may be separate species.
H. d. rufula. From Iberian Pen., s France, Corsica, Italy,
s Balkans, Greece and Cyprus, and from Morocco to w Algeria e through w,s
Turkey, Near East, ne Iraq, s Iran, n Afghanistan and from Transcaspia
e to Turkestan and e Kazakhstan to n Pakistan, nw India in the Himalayas,
1000-3300 m in Kashmir and Himachal Pradesh, and probably w China; sw Arabia;
locally in mts. in Sierra Leone and Cameroon; highlands from C. African
Rep., e Zaire, extreme se Sudan, s Ethiopia and n Somalia, s through Uganda,
Kenya and Tanzania to se Zaire, n Zambia and Malawi.
H. d. melanocrissus. Highlands of w,c Ethiopia.
H. d. daurica. From the Irtysh River area in Kazakhstan,
s Siberia from Altai e through Transbaicalia and L. Baikal region to Amurland
and Ussuriland, Korea, c.s Japan from c Honshu to Kyushu, and Izu Is.,
s through c,e China to India and se Tibet.
H. d. hyperythra. Sri Lanka.
Hirundo domicella WEST AFRICAN SWALLOW. Open country, towns.
Lowlands from Senegambia and Guinea e across s Mali, n Ghana, c Beni, s
Niger, Nigeria and n,c Cameroon to s Sudan (exc. extreme se) and extreme
cw Ethiopia. Often included in H. daurica -- which may be correct.
Dowsett and Dowsett-Lemaire (1993, p. 349) do not accept the separation
of domicella from daurica, noting that H. daurica has a wide range (see
above) and that other populations should be studied before such a conclusion
is drawn -- and that much of the area involved has not been explored.
This is one of many such situations which are products of differing species
concepts and lack of detailed evidence.
Hirundo striolata STRIATED SWALLOW. Open country, towns.
Lowlands to 1200 m from sw China, ne India and Taiwan, s through se Asia
to Sumatra (possibly), Java, Borneo, Lesser Sunda Is. e to Wetar and Timor,
and Philippines (except from Mindanao to Sulu Arch). Sometimes included
in H. daurica.
Hirundo spilodera group: H. perdita, preussi, rufigula,
spilodera. All occur in Africa. H. perdita glossy blue-black
above and throat; rump gray; buffy-orange breast band and crissum; white
belly; preussi back and crown blue-black; red spot behind eye; buffy rump
and below; shallow tail fork; rufigula like preussi without red spot and
darker rufous rump and underparts; spilodera similar, with red lores, spotted
buffy throat, rusty rump. Colonial nesters; gourd-shaped nests of
mud with short, tubular entrances, facing downward; often several hundred
or more nests clustered together; under bridges, eaves, rock faces, etc.
Eggs 1-4, usually 3, white with reddish, purple, gray markings.
Hirundo perdita RED SEA SWALLOW. Open country. Known
only from the type specimen found dead on the Red Sea coast of Sudan; presumably
breeding in the hills of ne Sudan and n Ethiopia. There have been
possible sight records in Ethiopia (Madge and Redman, 1989. Scopus 13:126-129).
Hirundo preussi PREUSS'S SWALLOW. Open country, towns.
From Guinea-Bissau (possibly), Sierra Leone and s Mali e to Ivory Coast,
Burkina Faso, n Ghana, Benin, Nigeria, Cameroon, Chad and ne Zaire.
Sometimes considered conspecific with rufigula, but they differ and are
allopatric.
Hirundo rufigula RED-THROATED SWALLOW. Open country, towns.
From Gabon, Congo River mouth and sc Zaire s to c Angola and nw Zambia.
Hirundo spilodera SOUTH AFRICAN SWALLOW. Open country, towns.
Sw Zimbabwe, s Zaire, Botswana, Namibia and S. Africa in Transvaal, Orange
Free State, w Natal and n,e Cape Province.
Hirundo pyrrhonota CLIFF SWALLOW. Open country, usually
near water, towns. From w,c Alaska, c Yukon, n Mackenzie, c Keewatin
and n Manitoba e across ec,se Canada to New Brunswick, Prince Edward I.
and Nova Scotia and s to sc,s Alaska, n Baja Calif., Oaxaca, Veracruz,
s Texas, sw Louisiana, n portions of Gulf states, w S. Carolina and s N.
Carolina; locally in s Florida in Lake Okeechobee area.
Often placed in Petrochelidon. Nests in colonies, often of 1000-3000
pairs; retort-shaped nest of mud pellets with tubular opening, plastered
on vertical surface under an overhanging rock, under eaves, bridges, culverts
or similar locations; lined with grass, feathers; built in 5-14 days; may
be repaired and reused. Other birds often use vacant nests, including
House Sparrow and Plain Titmouse. House Sparrow may usurp a nest.
Intraspecific brood parasitism apparently frequent; cowbird parasitism
rare. Eggs 3-6, white, creamy, or pinkish white with reddish-brown
spots.
Hirundo fulva group: Cave (H. fulva) and Chestnut-collared
(H. rufocollaris) swallows. The Cave Swallow occurs from the sw U.S.
to Ecuador and the Greater Antilles. It hybridizes in a few places
with H. rustica. Plumage similar to Cliff Swallow, but forehead rufous
and throat buffy. The Chestnut-collared Swallow occurs in a narrow
strip from sw Ecuador to nw Peru; similar to Cave Swallow but throat and
cheeks white; nuchal collar rufous. Nest of Cave Swallow is an open
cup of mud, plastered to an overhanging rock, culvert, bridge. Nest
of Chestnut-collared is an enclosed ball-shaped or retort-shaped structure
with a side entrance often extended as a short tube, much like that of
a Cliff Swallow; in colonies on cliffs or under eaves. The taxonomic
relationships among the Cliff, Cave and Rufous-collared swallows remain
unclear, and the occasional hybridization between Cave and Barn swallows
indicates close genetic relationship between the rustica group and the
fulva group.
Hirundo fulva CAVE SWALLOW. Open country, ravines, towns.
These two groups, which are not in contact, sometimes are treated as
separate species.
H. f. pelodoma. From s Arizona, se New Mexico and w,sc,se
Texas s in n Mexico to e Durango, Zacatecas, San Luis Potosí and
w Tamaulipas; rapidly expanding its range in recent years.
H. f. fulva. C Chiapas, Yucatán and Quintana Roo;
s Florida; Cuba, Isle of Pines, Hispaniola, Gonâve I., Tortue and
Vieques islands, and Î.-à-Vache, Jamaica and Puerto Rico.
Hirundo rufocollaris CHESTNUT-COLLARED SWALLOW. Open country,
ravines, towns. In extreme sw Ecuador in Guayaquil and Loja, and
w Peru s to Lima. Sometimes treated as a race of H. fulva.
Hirundo nigricans TREE MARTIN. Open woodland, towns.
Lesser Sunda Is. of Timor and Flores; Australia; possibly s New Guinea.
Has been placed in Cecropis.
Glossy blue-black above with whitish rump, rufous forehead; buffy-white
below; black tail with shallow notch. Nests in small colonies; nest
of grass and leaves in a hollow tree limb, the opening sometimes partly
closed with mud; several pairs may share a single tree-hollow; also nests
in holes in cliffs or buildings, or usurps nest of Fairy Martin.
Eggs 3-5, pinkish white flecked with pale rufous at larger end.
Hirundo ariel group: Streak-throated Swallow (H. fluvicola)
and Fairy Martin (H. ariel). The Streak-throated Swallow occurs in
s Asia from Afghanistan to India; the Fairy Martin in Australia and Tasmania.
H. fluvicola is glossy steel-blue above; crown dark rufous; rump brownish;
below buffy-white with dark streaks. H. ariel dull blue-black above;
crown rufous; rump and underparts white. Both species are colonial,
often with many nests run together in a single mass; nest gourd-shaped
of mud pellets with tubular entrances 5-30 cm long; colonies under bridges,
arches, eaves, overhanging rock faces, in caves, hollow trees, etc.
Nest chamber lined with grass, feathers; eggs 3-5, white with rufous or
yellowish spots.
Hirundo fluvicola STREAK-THROATED SWALLOW. Open country
near water, towns. Lowlands to 1000 m in ne Afghanistan, n Pakistan
and n,c,peninsular India w to Gujarat, c Rajasthan and Punjab, n to foothills
of the Himalayas, and e to w Nepal, Bihar and Orissa. Closely related
to ariel.
Hirundo ariel FAIRY MARTIN. Open woods near water, towns.
Australia except extreme n Queensland, extreme sw W. Australia. Ranges
to Timor and New Guinea.
Hirundo fuliginosa FOREST SWALLOW. Forest, towns.
Lowlands from Nigeria and s Cameroon s to Equatorial Guinea (Rio Muni)
and c Gabon.
12 cm. Entirely dark brown; crown black with bluish gloss; chin
and throat tinged with rufous; tail slightly forked. Nest a hemispherical
chamber of mud pellets with one or two short, tubular, openings; plastered
to rock surface of cave roof or in rock crevice in forest; chamber lined
with plant down; eggs 2-3, white or with few pale rufous spots; solitary
nester, but may be a cooperative breeder since four adults slept in one
nest.
Delichon: House-Martins Delichon is closely related to the
Petrochelidon complex in Hirundo and may be included in Hirundo.
Eurasia, nw Africa. Northern House-Martin (D. urbica) occurs across
Eurasia, nw Africa; glossy blue-black above; rump and underparts white;
tail short, notched. Tarsi feathered in all three species.
Asian House-Martin (D. dasypus) is often considered conspecific with urbica
and called D. u. cashmeriensis by some authors, but it is probably sympatric
with urbica in eastern Asia (n. Pakistan); dasypus is like urbica in plumage,
but is grayish-white below with black-tipped upper tail coverts.
Nepal House-Martin (D. nipalensis) like urbica, with a broken white collar
on hindneck; chin, throat and crissum black. All species colonial;
nest a hemispherical structure of mud pellets plastered to a vertical surface;
a small, rounded aperture at the top rim or just below; lining of grass,
feathers; 3-5 white eggs.
Delichon urbica NORTHERN HOUSE-MARTIN. Open country, towns.
From British Isles, Shetland Is. and n Scandinavia e across n Russia and
n Siberia to w Anadyrland, Sea of Okhotsk and Sakhalin, s to nw Africa
from Morocco e to n Libya, n Mediterranean region incl. most islands, Turkey,
Near East, n Iraq, Iran, Afghanistan, Turkestan and e Kazakhstan, but absent
from region between Caspian Sea e to Lake Balkhash; nw India in the Himalayas,
3000-4500 m of n Kashmir and Ladakh, w China in w,n Sinkiang, Mongolia
and Manchuria in Heilungkiang; small colonies sometimes breed in s Africa
(e.g., Botswana) apparently European birds that remain after migration.
Delichon dasypus ASIAN HOUSE-MARTIN. Open country, towns.
Mts. to 5000 m, mostly above 1500 m, in n Pakistan in Gilgit, n India from
Kashmir e to Arunachal Pradesh, s Tibet, c,se China, Korea, se Siberia
in Sakhalin, Kuril Is., Japan on Hokkaido, Honshu and Kyushu, and Taiwan.
Often included in urbica, but close approaches and probable sympatry at
several places in e Asia indicates allospecies.
Delichon nipalensis NEPAL HOUSE-MARTIN. Open country, towns.
Himalayas, 1800-4000 m from nc India to w Burma and n Vietnam in nw Tonkin.
Psalidoprocne: Sawwings. Africa. Some authors
recognize only five species with seven as subspecies of P. pristoptera.
Medium-sized swallows with rough outer edge of primaries in males due to
barbs being curved. Glossy black, blue or green, or unglossed brown;
underwing coverts often white or gray; tail deeply forked, except in nitens;
nest in a tunnel or on a ledge (fuliginosa); eggs 2-4, white.
Dowsett and Dowsett-Lemaire (1993, p. 349) question the recognition
of several populations as species and suggest that the P. pristoptera group
may consist only of nitens, pristoptera, obscura, albiceps and fuliginosa.
They acknowledge the possibility that "...the "species" P. pristoptera
does indeed hide more than one biologically distinct taxon, but detailed
study of various breeding populations ... is needed to determine any such
important distinctions." This can be said of many avian taxa.
Psalidoprocne nitens SQUARE-TAILED SAWWING. Forest.
From Sierra Leone e to s Cameroon and n,ne Zaire, and s to Congo River
mouth and c,ce Zaire. Blackish-brown with greenish gloss above and
below; tail not forked. Solitary nester in a tunnel, chamber lined
with lichens, moss.
Psalidoprocne fuliginosa MOUNTAIN SAWWING. Montane forest,
clearings. Mts. of sw Cameroon on Mt. Cameroon and Fernando Po Island
in the Gulf of Guinea; recorded in Nigeria. Dull brown above and
below; tail forked; solitary nester or in small groups; nest of lichens,
moss; on a ledge, in a hole, cave, inside building.
Psalidoprocne albiceps WHITE-HEADED SAWWING. Savanna, clearings.
S Sudan, Uganda, w Kenya, e Zaire, n,w,s Tanzania, Malawi, n Zambia and
ne Angola.
Psalidoprocne pristoptera group: C, se, ne Africa. The eight taxa
here treated as species are geographically allopatric and similar in plumages;
sometimes considered to be subspecies of pristoptera: P. pristoptera,
antinorii, oleaginea, mangbettorum, chalybea, petiti, orientalis, holomelas
(holomelas, not holomelaena, is correct). Plumages glossy purplish
to greenish-black or brownish with white, gray or brown underwing coverts.
Male tail deeply forked; female less so. Variations include depth
of tail fork; extent of white or gray underwing area. Solitary nesters
or in small groups; nest in a tunnel dug by both adults in a sandy cliff,
earth bank, stream bank; or in an abandoned bee-eater nest tunnel; nest
chamber lined with lichens and dry grass.
Psalidoprocne chalybea SHARI SAWWING. Moist woodland, forest.
In extreme se Nigeria, Cameroon (exc. sw), C. African Rep., and n,ce Zaire
(exc. extreme ne).
Psalidoprocne petiti PETIT'S SAWWING. Moist woodland, forest.
From se Nigeria and sw Cameroon s to Cabinda and lower Congo River.
Psalidoprocne mangbettorum MANGBETTU SAWWING. Moist woodland,
forest. In extreme ne Zaire and extreme sw Sudan.
Psalidoprocne oleaginea ETHIOPIAN SAWWING. Moist woodland,
forest. Sw Ethiopia.
Psalidoprocne pristoptera BLUE SAWWING. Moist woodland,
forest. Eritrea.
Psalidoprocne antinorii BROWN SAWWING. Moist woodland, forest.
Highlands of wc Ethiopia (=blanfordi) and c,s,ne Ethiopia (= antinorii).
Psalidoprocne orientalis EASTERN SAWWING. Moist woodland,
forest. From sw Gabon, Cabinda, s,se Zaire and s,ne Tanzania (incl.
Usambara Mts.), s to sw,c,ne Angola, Zambia, Malawi, e Zimbabwe and adj.
w Mozambique.
Psalidoprocne holomelas BLACK SAWWING. Moist woodland,
forest. Highlands in ce Zaire, Uganda, Rwanda, Burundi, w,c Kenya
and ne,ce Tanzania s to Uluguru and Usambara mts.; ne Namibia, nw Botswana,
se Zimbabwe, s Mozambique, e,s S. Africa in e Transvaal, Natal, s,e Cape
Province.
Psalidoprocne obscura FANTI SAWWING. Ec Africa. Moist
woodland, forest. From Senegambia e to e Nigeria, sw Cameroon and
C. African Rep. Male blackish-brown with green sheen; crown and throat
white; black eyeline; female same but with variable white crown markings;
tail forked. Solitary nester in a tunnel in a dry site; nest of grass,
lichens, feathers.
Family REGULIDAE: Kinglets,
Goldcrest, Flamecrest, Firecrest. Coniferous and mixed forests in
N.A. and Eurasia; Taiwan; Canary Is.
9 cm. Small, greenish above, whitish to yellowish below;
white wing bars; differ mainly in crown markings: Ruby-crowned Kinglet
(R. calendula), w, n N.A., red crown patch; white eyering. Goldcrest
(R. regulus), Eurasia, male crown with orange center stripe, bordered black;
female crown center yellow. Canary Is. Kinglet (R. teneriffae), similar
to R. regulus. Flamecrest (R. goodfellowi), mts. of Taiwan, crown
black with concealed orange patch, white superciliary, brown ear region,
yellow rump, flanks. Firecrest (R.ignicapillus) c, s Europe, male,
center of crown red, bordered black, wide white superciliary, female crown
yellow center. Golden-crowned Kinglet (R. satrapa), male like calendula
with crown center red, bordered by yellow and black stripes to white superciliary.
Feed on insects gleaned from foliage, often hover to take prey from leaves.
Nest a bulky purse-like structure of moss suspended from, or attached to,
twigs or branchlets; eggs 5-10, white or buffy, usually with brown spots.
Regulus calendula RUBY-CROWNED KINGLET. Coniferous forest,
mixed woods, muskeg. From nw,nc Alaska, c Yukon, nw,s Mackenzie,
n,c Alberta and n,c Saskatchewan e across c Canada to Labrador and Newfoundland,
s in mts. to s Calif., c,s Arizona, sc New Mexico and ec Colorado, and
e of the Rockies to c,ne Minnesota, ne Wisconsin, n Michigan, s Ontario,
n New York, n Maine and Nova Scotia; Guadalupe I. off Baja Calif.
Regulus regulus GOLDCREST. Coniferous and mixed forest.
From British Isles and Scandinavia e across nw,c Russia to sw Siberia (
Altai, Lake Baikal) and s to Azores, n Mediterranean region, Corsica, Sardinia
and Sicily, Turkey, Syria, n Iran and Crimea, Caucasus; Himalayas to 4400
m in e Turkestan, ne Afghanistan, n Pakistan, n India from Kashmir e to
Arunachal Pradesh, and c China, Manchuria, se Siberia to Sakhalin, s Kuril
Is., and Japan.
Regulus teneriffae CANARY ISLANDS KINGLET. Coniferous and
mixed forest. Canary Islands. Sometimes included in ignicapillus
or in regulus.
Regulus goodfellowi FLAMECREST. Coniferous and mixed forest.
Mts. above 2000 m of Taiwan.
Regulus ignicapillus FIRECREST. Mixed or coniferous forest.
From s England and continental Europe e to c Poland and s Russia, Carpathian
Mts., Crimea, w Caucasus, s to Madeira, Morocco to n Tunisia, n Mediterranean
region incl. most islands (except Crete and Cyprus), and Turkey.
Regulus satrapa GOLDEN-CROWNED KINGLET. Coniferous forest,
woods. From s Alaska w to base of Alaska Pen., s Yukon, n Alberta
and Saskatchewan e across s Canada to Newfoundland and Nova Scotia, s in
mts. to s,e Calif., w,ne Nevada, s Utah, c,se Arizona, sc New Mexico, in
highlands of Mexico to w Guatemala, and e of the Rockies to n,ec Minnesota,
n Illinois, c Indiana, n Ohio, s Ontario, New York, and c New England,
s in the Appalachians to e Tennessee and w N. Carolina.
Family PYCNONOTIDAE: Finchbills,
Bulbuls, Greenbuls, Bristlebills, Nicators. Old World tropics, a
few species in temperate areas of se Asia. Sexes alike; rictal bristles
and long bristles extending from nape; lush rump plumage. Arboreal
or in scrub, thickets. Food: fruits, seeds, insects, nectar, pollen.
Nest an open cup of leaves and plant fibers in a tree; eggs whitish or
pinkish with reddish-brown markings; variable. Female builds nest,
incubates.
Spizixos: Finchbills. S Asia.
20 cm. Bills heavy, finchlike; live in scrub, second-growth.
Crested Finchbill (S. canifrons), head blackish with long crest; forehead,
throat, cheeks gray; olive-green above; yellowish-green below. Collared
Finchbill (S. semitorques); like Crested with shorter crest; head, throat
blackish; nape gray; white collar and lores; cheeks white-streaked.
Nest a shallow cup of curly vine tendrils, twigs, lined with grass, ferns;
in low bushes or small trees; 2-4 dull pink eggs with reddish speckles.
Some are communal breeders.
Spizixos canifrons CRESTED FINCHBILL. Scrub, second growth,
forest. Hills, 1000-2500 m in e India, se Tibet, sw China, nw Thailand,
n Laos and n Vietnam in nw Tonkin.
Spizixos semitorques COLLARED FINCHBILL. S China, Taiwan.
Scrub, second growth, towns. C,s China, Taiwan and n Vietnam in nw
Tonkin.
Pycnonotus: Six species occur in Africa, 36 in s, se Asia;
most are common, vocal birds of second growth, gardens, woods, forests.
Most 10-18, up to 29 cm. Plumages variable, some crested; most
dull-colored, some with red, yellow, orange, black or white areas.
Pycnonotus zeylonicus STRAW-HEADED BULBUL. Riparian growth,
scrub. Lowlands to 1600 m in the Malay Pen., Sumatra, Nias, Java
and Borneo.
Pycnonotus striatus STRIPED BULBUL. Forest. Himalayas,
1200-2750 m in ne India, sw China, Burma, nw Thailand and n Vietnam.
Pycnonotus leucogrammicus CREAM-STRIPED BULBUL. Forest.
Highlands, 800-1900 m of Sumatra.
Pycnonotus tympanistrigus SPOT-NECKED BULBUL. Forest.
Foothills, 300-900 m of Sumatra.
Pycnonotus melanoleucos BLACK-AND-WHITE BULBUL. Forest,
second growth. Lowlands to 1800 m in peninsular Thailand, Malaya,
Sumatra incl. Siberut, and Borneo.
Pycnonotus priocephalus GREY-HEADED BULBUL. Forest.
Lowlands to 1800 m of peninsular sw India in the western Ghats from extreme
s Maharashtra, Goa and w Mysore s to Kerala.
Pycnonotus atriceps BLACK-HEADED BULBUL. Forest, edge,
scrub, second growth. Lowlands to 2450 m in e India, sw China, Burma,
Thailand, Cambodia, Laos, n Vietnam, Malaya, Andaman Is., Sumatra incl.
Simeulue, Nias, Mentawai Is., Bangka and Belitung; Borneo incl. Maratua
Is.; Java incl. Bawean I.; Bali; Calamian Is. and Palawan in the sw Philippines.
Pycnonotus melanicterus BLACK-CRESTED BULBUL. Forest, second
growth, scrub.
Some allopatric populations may be separate species.
P. m. flaviventris. Lowlands to 2400 m in India, sw China
and n Burma.
P. m. gularis. Peninsular India in the W. Ghats from w
Mysore s to Kerala and Tamil Nadu.
P. m. melanicterus. Sri Lanka.
P. m. dispar. Se Asia except n Burma, Sumatra, Java and
highlands of n Borneo.
Pycnonotus squamatus SCALY-BREASTED BULBUL. Forest.
Lowlands to 1200 m in Malay Pen., Sumatra, w,c Java and Borneo.
Pycnonotus cyaniventris GREY-BELLIED BULBUL. Forest, open
woods. Lowlands to 1000 m in Malay Pen., Sumatra incl. Sipura, and
Borneo.
Pycnonotus jocosus RED-WHISKERED BULBUL. Forest edge, second
growth, towns. Lowlands to 1500 m in India, Burma, se China, se Asia
(except s Malaya), Andaman Is. Intro. Mauritius, e,se Australia,
Nicobar Is., Hawaiian Is. on Oahu and Florida.
Pycnonotus xanthorrhous BROWN-BREASTED BULBUL. Scrub, second
growth. Himalayas, 1200-4300 m, rarely down to 600 m, in c,e,s China,
se Tibet, ne,e,s Burma, nw Thailand, n Laos and n Vietnam.
Pycnonotus sinensis LIGHT-VENTED BULBUL. Second growth,
farms. Lowlands in s,se China, Hainan I., n,e Vietnam, Taiwan (exc.
s) and s Ryukyu Is.
Pycnonotus taivanus STYAN'S BULBUL. Second growth, farms.
Coastal lowlands of e,s Taiwan.
Pycnonotus cafer group: Of the following 11 species, six occur
in Africa, five in Asia. The six African species listed below are barbatus,
somaliensis, dodsoni, tricolor, nigricans, capensis. The first four
are often considered to be conspecific as the "barbatus group". They
are similar in plumages, have complex distributions and often hybridize,
but no two forms hybridize in all areas of overlap. The situation
is complex and not easily reduced to a simple set of names. The members
of the barbatus group have brown heads, breasts, backs and tails; brownish
or whitish below; crissum yellow or white.
"The allegation that P. somaliensis, dodsoni and tricolor occur in Ethiopia
(plus barbatus) without interbreeding is not without question; but before
accepting such, the East African List Committee would want to see more
evidence of their "different ecological requirements". For the time
being we consider them all conspecific with barbatus. (D. A. Turner,
pers. comm.). This statement refers to the text under P. barbatus
in Sibley and Monroe 1990:583. Dowsett and Dowsett-Lemaire (1993,
p. 352) are critical of the Sibley and Monroe (1990) treatment (based on
Hall and Moreau, 1970) which recognized the four species listed above --
but they note that "No critical collection of data for analysis has been
made in any of these regions, and our knowledge remains as presented by
Hall and Moreau (1970)." However, they accept only two species (barbatus,
nigricans) and note that their field experience "suggests that in most
parts of Africa there is but one". Clearly, the situation is complex
and information is conflicting and incomplete. Dowsett and Dowsett-Lemaire
(1993) would include somaliensis, dodsoni and tricolor as subspecies of
barbatus. They may be right, but they also suggest that "the matter
does require further investigation ...." This is but one among many
such problems that reflect ignorance of the details in nature, as well
as different species concepts. Whether viewed as one, two, three
or four species -- members of the barbatus group occur in every country
in continental Africa.
Pycnonotus barbatus GARDEN BULBUL. Woods, forest, second
growth, towns, usually near water.
P. b. barbatus. From Morocco e to n Tunisia; from Senegambia,
Sierra Leone and Liberia e through s Mali, s Niger and s Chad to w Sudan,
and s in wc Africa to Gabon and lower Congo River basin.
P. b. arsinoe. From Nile Valley of Egypt and n,c Sudan
e to w,c Ethiopia.
Pycnonotus somaliensis SOMALI BULBUL. Bush, woods, usually
in arid regions. C,e Ethiopia and nw Somalia.
This species and the next two (three ?) species may be conspecific with
barbatus; they are similar in plumages, have complex distributions and
often hybridize.
Pycnonotus dodsoni DODSON'S BULBUL. Bush, woods, in arid
regions. C,se Ethiopia, Somalia and Kenya (exc. w,sc).
Pycnonotus tricolor DARK-CAPPED BULBUL. Woods, forest edge,
second growth, towns, usually near water. From e,s Cameroon, s Chad,
C. African Rep., c,s Sudan and c,sc Ethiopia s to s Angola, ne Namibia,
n,e Botswana and e S. Africa in Transvaal, Natal, e Orange Free State,
e Cape Province.
Pycnonotus nigricans BLACK-FRONTED BULBUL. Dry scrub, usually
near water. Arid regions from s Angola, Botswana and sw Zimbabwe
s to Namibia and interior S. Africa e to Transvaal and Lesotho.
This species is viewed as a member of the cafer group; it should be
included in the African barbatus group because it hybridizes with barbatus
in Angola, Botswana and Transvaal, but they also occur in sympatry along
range boundaries without interbreeding.
Pycnonotus capensis CAPE BULBUL. Dense scrub, riparian bushes.
Sw,s,e Cape Province in S. Africa.
Like nigricans, capensis is closely related to the members of the barbatus
group. P. capensis overlaps with nigricans and barbatus in s Cape
Province without interbreeding.
The five Asian species of the cafer group are xanthopygos, leucotis,
leucogenys, cafer, aurigaster.
Pycnonotus xanthopygos WHITE-SPECTACLED BULBUL. Woods,
towns, second growth. C,s Turkey, Near East, Sinai Pen. of Egypt
and Arabia, except cn. Has hybridized with barbatus in Egypt, may
be conspecific.
Pycnonotus leucotis WHITE-EARED BULBUL. Dry scrub, farms,
woods, mangroves, towns. Lowlands to 1800 m from e,se Iraq, s Iran
and cn Arabia e to s Afghanistan, Pakistan and w India n to Himalayan foothills
and s to n Maharashtra and n Madhya Pradesh. Closely related to leucogenys,
may be conspecific. They occasionally hybridize in w India (Sibley
and Short 1959. Ibis 101:177-182).
Pycnonotus leucogenys HIMALAYAN BULBUL. Dry scrub, farms,
woods. Foothills and mts., 600-2750 m from e Afghanistan e in Himalayas
to ne India in Assam. Hybridizes with cafer in ne Pakistan, but occurs
widely with cafer in w India without interbreeding. Hybrids occur
where one of the species is rare.
Pycnonotus cafer RED-VENTED BULBUL. Scrub, second growth,
woods, towns. Lowlands to 1800 m in e Pakistan, India, Sri Lanka,
Burma and sw China. Intro. n Arabia, Polynesia, Oahu. Hybridizes
with leucogenys, see above.
Pycnonotus aurigaster SOOTY-HEADED BULBUL. Scrub, second
growth, woods, towns. Lowlands to 1800 m in s China, s,e Burma, Thailand
(exc. pen.), Indochina, Java and Bali (possibly introduced). Intro.
Sumatra and Sulawesi. Hybridizes with cafer in extreme e India.
P. aurigaster consists of several variable populations possibly reflecting
a hybrid origin.
Pycnonotus eutilotis PUFF-BACKED BULBUL. Second growth,
scrub, forest. Lowlands in Malay Pen., Sumatra and Bangka, and Borneo.
Pycnonotus nieuwenhuisii BLUE-WATTLED BULBUL. Presumably
forest or second growth. Known from two specimens from two localities;
nw Sumatra at 700 m; ne Borneo at 600 m.
Pycnonotus urostictus YELLOW-WATTLED BULBUL. Forest edge,
open woods, second growth. Lowlands to 1000 m of the Philippines
on Luzon, Polillo Is., Catanduanes, Samar, Leyte, Bohol, Panaon, Dinagat,
Siargao, Mindanao and Basilan.
Pycnonotus bimaculatus ORANGE-SPOTTED BULBUL. Second growth,
scrub. Highlands, 800-3000 m, in Sumatra, Java and Bali.
Pycnonotus finlaysoni STRIPE-THROATED BULBUL. Second growth,
scrub. Lowlands to 1000 m in sw China, Burma, Thailand, Indochina
and Malaya.
Pycnonotus xantholaemus YELLOW-THROATED BULBUL. Thorn scrub,
open forest. Hills, 600-1200 m of s India.
Pycnonotus penicillatus YELLOW-EARED BULBUL. Open forest,
woods. Hills above 900 m of Sri Lanka.
Pycnonotus flavescens FLAVESCENT BULBUL. Open forest, edge,
second growth, mossy forest. Hills and mts., 500-3500 m in e India,
Burma, sw China, nw Thailand, n,s Laos and n,c Vietnam; mts. 900-3500 m
of n Borneo from Mt. Mulu and Mt. Murud e to Mt. Kinabalu.
Pycnonotus luteolus WHITE-BROWED BULBUL. Dry scrub, towns,
second growth. Lowlands to 1000 m of s India on coasts n to Gujarat,
s Madhya Pradesh ans s W. Bengal; Sri Lanka.
Pycnonotus goiavier YELLOW-VENTED BULBUL. Second growth,
scrub, towns, mangroves. Lowlands, mostly coastal, up to 1500 m from
s Burma in s Tenasserim, e to s Vietnam, interior Cambodia and mts. of
Malaya; Sumatra incl. Riau and Lingga arch., Bangka and Belitung; Borneo
incl. Maratua Is.; Java incl. Karimunjawa Is.; Bali, Lombok and the Philippines
incl. the Sulu Arch. Sulawesi where probably introduced.
Pycnonotus plumosus OLIVE-WINGED BULBUL. Forest edge, second
growth, scrub. Lowlands to 600 m in Malay Pen. incl. Tinggi I., Sumatra
and most adj. small is., Borneo incl. Anambas, Balembangan, Banggi and
Mallawli islands; Java incl. Bawean, Bali; sw Philippines on Calamian Is.,
Palawan and Cagayan Sulu.
Pycnonotus blanfordi STREAK-EARED BULBUL. Second growth,
scrub, towns. Lowlands to 1000 m of se Asia (except ne Burma, n Vietnam
and s Malaya).
Pycnonotus simplex CREAM-VENTED BULBUL. Forest. Lowlands
to 1300 m in s peninsular Thailand, Malaya, Sumatra and Nias, Batu Is.,
Bangka, Belitung, Riau and Lingga arch.; Borneo and Anambas, Natuna and
Balembangan is., and Java.
Pycnonotus brunneus RED-EYED BULBUL. Forest, second growth,
scrub, edge. Lowlands to 1000 m in Malay Pen., Tioman I., Sumatra
and Nias, Banyak and Batu is., probably Bangka, and Borneo, Anambas, Karimata,
Banggi and Larot islands.
Pycnonotus erythropthalmos SPECTACLED BULBUL. Forest.
Lowlands to 1000 m in Malay Pen., Sumatra, Nias, Banyak and Batu is., Lingga
Arch., Bangka and Belitung; and Borneo.
Andropadus: Greenbuls. Confined to the forests of Africa;
some species occupy small areas; most live in the forested belt across
central Africa. Plumages are mainly uniform greenish, olive or brown, paler
below with yellowish abdomen; some with gray heads; none crested.
Sixteen species are listed here, but only 11 are recognized by some authors,
as noted below.
D. A. Turner (pers. comm.) offers the following comments: "Moyer and
Stuart (in prep.) will present evidence for the lumping of the East African
A. masukuensis with the West African nominate tephrolaemus. Since
tephrolaemus is the earlier name, the correct name for Shelley's Bulbul
would become A. tephrolaemus. At the same time East African montane
greenbuls would change from tephrolaemus to nigriceps. Within this
complex the East African List Committee agree that the s Tanzanian form
chlorigula is worthy of specific status, but the status of fusciceps and
other s Tanzanian forms is currently unclear. Meanwhile, the East
African List Committee do not consider kakamegae worthy of specific status."
It seems best to await the publication of the Moyer and Stuart study before
reorganizing the following arrangement.
Andropadus montanus CAMEROON GREENBUL. Forest. Mts.,
550-1800 m in Togo, possibly Nigeria, and Cameroon.
Andropadus masukuensis SHELLEY'S GREENBUL. Forest.
Mts., 900-2300 m in s Tanzania and n Malawi.
Andropadus kakamegae KAKAMEGA GREENBUL. Forest. Mts.,
900-2300 m in ne,ce Zaire, w Kenya and w Tanzania. Often treated as a subspecies
of masukuensis; the two are allopatric.
Andropadus virens LITTLE GREENBUL. Open forest, edge, second
growth. Lowlands and mts. from Senegambia, Sierra Leone and Liberia
e to Cameroon, Fernando Po I., C. African Rep., n,e Zaire, extreme s Sudan,
Uganda, w,se Kenya and Tanzania, Mafia I. and Zanzibar, s to n,ne Angola,
n Zambia, Malawi and n Mozambique.
Andropadus hallae HALL'S GREENBUL, known from one specimen from
e Zaire in the range of virens seems to be a melanistic individual of virens.
It is not recognized as a species by the East African List Committee (D.
A. Turner, pers. comm.).
Andropadus gracilis GREY GREENBUL. Forest. Lowlands
from Sierra Leone, se Guinea and Liberia e to s Cameroon, C. African Rep.,
n,ne Zaire, Uganda and w Kenya, s to nw Angola and sw,sc,ce Zaire s to
Lake Tanganyika.
Andropadus ansorgei ANSORGE'S GREENBUL. Forest. Lowlands
in a patchy distribution from Sierra Leone and Liberia e to s Cameroon,
n,ne Zaire, w Uganda and w Kenya, s to n Angola and sw,sc,ce Zaire.
Andropadus curvirostris PLAIN GREENBUL. Forest. Lowlands
from Sierra Leone and Liberia e to s Cameroon, Fernando Po I., s C. African
Rep., n,ne Zaire, s Sudan, Uganda and w Kenya, and s to nw Angola and sw,sc,ce
Zaire.
Andropadus gracilirostris SLENDER-BILLED GREENBUL. Forest.
Lowlands to 2100 m from Sierra Leone, se Guinea and Liberia e to s Cameroon,
Fernando Po I., sw C. African Rep., n,ne Zaire, s Sudan, Uganda and w Kenya,
s to n Angola, se,cs,se Zaire and w Tanzania.
Andropadus importunus SOMBRE GREENBUL. Some isolated populations
may be separate species.
A. i. insularis. Dense brush, scrub. Lowlands from s Somalia,
e Kenya, Tanzania and Zanzibar, s to se Zambia, Malawi and n Mozambique.
A. i. fricki. Humid forest. Highlands of c Kenya.
A. i. importunus. Humid forest. Zimbabwe, sc,s Mozambique
and se,s S. Africa in n,e Transvaal, Swaziland, Natal and sw,s,e Cape Province.
Andropadus latirostris YELLOW-WHISKERED GREENBUL. Humid
forest. Lowlands in the west to mts. in the east-central of w,c Africa
from Sierra Leone, se Guinea and Liberia e to s Cameroon, Fernando Po I.,
n,ne Zaire, s Sudan, Uganda and w,nc Kenya, s to nw Angola, sw,sc,ec Zaire
s to Lake Tanganyika and w Tanzania.
Andropadus tephrolaemus GREY-THROATED GREENBUL. Forest.
Mts. in se Nigeria, sw Cameroon and Bioko I. A. tephrolaemus, nigriceps
and chlorigula are sometimes considered conspecific, but their relationships
are unclear; the first two differ vocally.
Andropadus nigriceps MOUNTAIN GREENBUL. Forest. Locally
in mts. in s Kenya, w,ce,sw Tanzania, extreme e Zambia, Malawi and ne Mozambique.
Differs vocally from tephrolaemus and may be more closely related to masukuensis.
Andropadus milanjensis STRIPE-CHEEKED GREENBUL. Forest.
The East African List Committee consider olivaceiceps to be a subspecies
of milanjensis. Vocalizations are identical from Zimbabwe n to Kenya.
A. m. olivaceiceps OLIVE-HEADED GREENBUL. Forest.
Locally in mts. in se Kenya, ne,ce,sc,sw Tanzania, Malawi and extreme nw
Mozambique.
A. m. milanjensis. Locally in mts. in extreme e Zimbabwe
and cw Mozambique.
Calyptocichla serina GOLDEN GREENBUL. Second growth forest,
forest canopy. Locally from Sierra Leone e to s Cameroon, Bioko I.,
sw C. African Rep., and nc,ne,ce Zaire, s to Gabon and Congo River mouth.
Nest of dry leaves, plant fibers; in a fork at the end of a high branch
in crown of tall tree.
Baeopogon: Honeyguide Greenbuls. Short, broad bill;
olive-green above, gray or buff below; tail short with three outer rectrices
all, or partially, white, thus resembling honeyguides.
Baeopogon indicator HONEYGUIDE GREENBUL. Forest edge, second
growth, riparian vegetation. From Sierra Leone, se Guinea and Liberia
e to Cameroon, s C. African Rep., n,ne Zaire, s Sudan, Uganda and w Kenya,
and s to nw Angola, extreme nw Zambia and cs,se,ce Zaire.
Baeopogon clamans WHITE-TAILED GREENBUL. Locally in Cameroon,
Rio Muni, w Gabon and ne Zaire. These are disjunct populations.
Ixonotus guttatus SPOTTED GREENBUL. Forest, edge.
From Liberia e to s Cameroon, n,ne Zaire and w Uganda, s to sw,sc Zaire
and nw Tanzania.
Olive-brown above with white or yellowish spots on lower back and tail;
whitish to yellowish below; three outer rectrices white. Cooperative
breeder, 4-6 birds feed young in nest and after fledging.
Chlorocichla: Greenbuls. Africa, in thickets, scrub, forest
edge, clearings; often in noisy groups. Long, stout bills; long rictal
bristles; tail rounded or slightly graduated; throat white or yellow.
Chlorocichla simplex SIMPLE GREENBUL. Forest, second growth.
From Sierra Leone, se Guinea and Liberia e to s Cameroon, s C. African
Rep., n,ne Zaire and extreme s Sudan, s to nw,ne Angola and cs,c Zaire.
Nest a small cup of twigs, plant fibers, cobwebs. Eggs 1-2, glossy
white to beige, pale olive, grayish; with reddish spots.
Chlorocichla flavicollis YELLOW-THROATED GREENBUL. Forest,
thickets. From Senegambia, Guinea and Sierra Leone e to Cameroon,
C. African Rep., se Sudan, n,ne Zaire, Uganda and w Kenya, s to c,se Angola,
w,c,n Zambia, n Malawi and w Tanzania. Geographically variable. Probable
cooperative breeder. Nest, eggs, similar to simplex.
Chlorocichla falkensteini YELLOW-NECKED GREENBUL. Forest
edge, second growth. From s Cameroon, s C. African Rep., and Rio
Muni s to sw Zaire and w Angola.
Chlorocichla flaviventris YELLOW-BELLIED GREENBUL. Forest undergrowth,
thickets, riparian growth. From w,s Angola, Zambia, se Zaire, Tanzania,
c,se Kenya and ne S. Africa in n,e Transvaal, Swaziland and Natal. Nest
cup of twigs, grass, etc.; both sexes build; 2 eggs, variable.
Chlorocichla laetissima JOYFUL GREENBUL. Dense forest thickets,
riparian forest. Mts., 1150-1700 m in ne,ce,se Zaire, extreme se
Sudan, Uganda, e Zaire, nw Kenya and extreme cn Zambia.
Chlorocichla prigoginei PRIGOGINE'S GREENBUL. Forest.
Locally in mts., 1200-1650 m, of ne Zaire, local and rare. A recently
described sibling species of laetissima.
Thescelocichla leucopleura SWAMP GREENBUL. Swampy areas
with palms, Pandanus forest, cultivation. From Sierra Leone, se Guinea
and Liberia e to Cameroon, C. African Rep., n,ne Zaire and w Uganda, s
to sw,cs,c Zaire.
Brownish-olive above; chin and throat whitish; breast olive with white
shaft streaks; whitish to yellow below; 4 outer rectrices with large white
tips. Cooperative breeder.
Pyrrhurus scandens LEAF-LOVE. Often placed in Phyllastrephus.
Swampy forest, thickets, open forest in savanna. Lowlands from Senegambia,
Guinea-Bissau and Sierra Leone e to c Cameroon, C. African Rep., n,ne Zaire,
extreme s Sudan and w Uganda, s to cw,cs,se Zaire and extreme w Tanzania.
Nest a small cup of leaves, grass, cobwebs; slung between twigs over a
stream. Eggs 2-3, whitish, buffy with brown marks.
Phyllastrephus: Greenbuls, Brownbuls. Forest undergrowth
in Africa and Madagascar; the five Madagascar species have been assigned
to Bernieria or Oxylabes and sometimes placed in the Timaliinae (babblers).
Plumages mainly brown or olive-green, some species yellow below; tail graduated
or rounded, often rufescent. Sexes differ in size.
The taxonomy of the species of Phyllastrephus is uncertain. Plumages
are so similar that they provide little information about relationships
and there are no molecular data to provide objective evidence.
Phyllastrephus cabanisi CABANIS'S GREENBUL. Forest. Highlands
above 1200 m from s Sudan, Uganda, w Kenya, w Tanzania and e Zaire to n
Zambia and w to n,c Angola.
P. cabanisi, fischeri and placidus are allopatric taxa sometimes treated
as conspecific; fischeri is vocally and morphologically distinct from the
other two. P. poensis (see below) may be a subspecies of cabanisi,
or it may be closest to baumanni.
Phyllastrephus fischeri FISCHER'S GREENBUL. Coastal lowland
forest and scrub. Lowlands from se Somalia and e Kenya to coastal
ne Mozambique. Plumage similar to that of cabanisi.
Phyllastrephus placidus PLACID GREENBUL. Forest undergrowth,
riparian scrub. Highlands of e Africa from w,c Kenya e of Rift Valley,
n,e,s Tanzania to n Zambia, Malawi and s to ne Mozambique.
The East African List Committee consider placidus conspecific with cabanisi.
The form sucosus is intermediate between the two, but vocally indistinguishable
(D. A. Turner, pers. comm.).
Phyllastrephus terrestris TERRESTRIAL BROWNBUL. Mainly lowland
riverine bush, dry thickets in savanna or acacia. From s Angola,
ne Namibia, Zambia, extreme se Zaire, s,e,ne Tanzania, c,se Kenya and s
Somalia s to n,e Botswana, n,e Zimbabwe and e,s S. African in n,e Transvaal,
Swaziland, Natal and s,e Cape Province. Nest a flimsy cup of twigs, stems
on a branch or slung between twigs, near ground.
Phyllastrephus strepitans NORTHERN BROWNBUL. Arid scrub,
thornbush, woods. From n Uganda, extreme se Sudan, sc,s Ethiopia
and s Somalia s through Kenya and e Tanzania to Mozambique border.
Similar to terrestris.
Phyllastrephus fulviventris PALE-OLIVE GREENBUL. Riparian
undergrowth. Sw Zaire, Cabinda and w Angola.
Phyllastrephus cerviniventris GREY-OLIVE GREENBUL. Riparian
thickets, undergrowth, forest. Nc,se Kenya, e,s Tanzania, Zambia
(exc.sw), Malawi and n,c Mozambique.
Phyllastrephus baumanni BAUMANN'S OLIVE-GREENBUL. Forest
undergrowth. From n,e Sierra Leone and Liberia e to sw,cs Nigeria.
Phyllastrephus poensis CAMEROON OLIVE-GREENBUL. Humid forest.
Mts. above 1400 m in se Nigeria, Cameroon and Fernando Po I. Closest
relatives uncertain - baumanni or cabanisi?
Phyllastrephus hypochloris TORO OLIVE-GREENBUL. Forest,
undergrowth. Lowlands and mts. in ne Zaire, extrene se Sudan, Uganda
and w Kenya.
Phyllastrephus lorenzi SASSI'S GREENBUL. Forest. Known
only from a few localities in ne,ce Zaire; patchy distribution, rare.
Phyllastrephus poliocephalus GREY-HEADED GREENBUL. Humid
forest. Mts. in Nigeria and sw Cameroon.
Phyllastrephus flavostriatus YELLOW-STREAKED GREENBUL.
Forest. Mts. and, in s, lowlands. Patchy distribution in from
ne,ce Zaire s to Lake Tanganyika and w Tanzania, sw,c Uganda and extreme
se Kenya s through c,e,coastal Tanzania, s Malawi, e Zimbabwe and Mozambique
to e S. Africa in Transvaal, Natal and extreme e Cape Prov. P. alfredi
may be a subspecies of flavostriatus, and is so treated by some authors.
Phyllastrephus alfredi SHARPE'S GREENBUL. Mt. rain forest.
Patchy distribution in extreme sw Tanzania and n Malawi. This range
is between the ranges of subspecies of flavostriatus. General habits
and voice like flavostriatus.
The ranges of the subspecies of flavostriatus and that of alfredi
are allopatric, each occupying a disjunct area of mountainous terrain.
Thus, the definition of alfredi as a separate species is based on its slightly
different plumage coloration. There is no objective way to decide
whether alfredi is a race of flavostriatus or a separate species; either
treatment may be defended and both are reasonable. The East African
List Committee consider alfredi conspecific with flavostriatus (D. A. Turner,
pers. comm.).
Phyllastrephus debilis TINY GREENBUL. Coastal scrub, lowland
forest, riparian forest. Se Kenya, coastal to inland ne, se Tanzania,
incl. foothills of Uluguru Mts., e Zimbabwe and c,s Mozambique; ne Tanzania
in the Usambara Mts. Smallest African bulbul. Nest a small, neat
cup of plant fibers, grass, etc. bound with cobwebs in a fork or outer
branches. Eggs 2, light blue with brownish spots.
Phyllastrephus albigularis WHITE-THROATED GREENBUL. Lowland
forest. From Sierra Leone and Liberia e to s Cameroon, s C. African
Rep., n,ne Zaire, extreme s Sudan, Uganda and extreme w Kenya, s to nw
Angola and sc,ec Zaire.
Phyllastrephus icterinus ICTERINE GREENBUL. Forest undergrowth.
Lowlands of w,c Africa from Sierra Leone e to s Cameroon, Fernando Po I.,
sw C. African Rep., n,e Zaire and extreme w Uganda, s to Gabon and sc,ce
Zaire.
Phyllastrephus leucolepis LIBERIAN GREENBUL. Humid forest.
Known only from two patches of rain forest in se Liberia.
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