Rollandia rolland WHITE-TUFTED GREBE.
Lakes, streams, ponds, marshes, bays. From nw Peru, c,s Bolivia,
Paraguay and s Brazil s to Rio Grande do Sul. Falkland Islands.
Rollandia microptera SHORT-WINGED GREBE.
Shallows of Andean lakes. Andes in the Lake Titicaca basin of s Peru
and w Bolivia.
Tachybaptus ruficollis LITTLE GREBE.
Marshes, ponds, lakes. From British Isles, s Sweden, c,s Russia n
to a line from Lithuania s to Caucasus and n Caspian Sea, s Russia n to
Caspian and Aral seas, and e to Lake Balkash, China incl. Hainan I., Tibet,
Taiwan, Japan and Kuril Is., s to n Africa locally from Morocco e to lower
Nile River; Mediterranean area incl. most islands; Near and Middle East,
Arabia incl. Yemen, Pakistan, India, Sri Lanka, se Asia, Indonesia in the
Greater and Lesser Sunda islands, Moluccas, Kai Is., Philippines incl.
Palawan; n New Guinea; Admiralty Is., Bougainville Island in the n Solomon
Is. Africa s of the Sahara from sw Mauritania, Senegal, Sierra Leone,
Liberia, Ivory Coast, Ghana, Togo, Nigeria, s Mali, Burkina Faso, s Niger,
Cameroon, Central Afr. Rep., s Chad and s Sudan e to Ethiopia, Eritrea,
Djibouti and Somalia, s locally through Gabon, Congo, Zaire, Rwanda, Burundi,
Uganda, Kenya, Tanzania, Angola, Zambia, Malawi, Mozambique, Namibia and
Botswana to s S. Africa; Madagascar; Comoro Islands.
Tachybaptus novaehollandiae AUSTRALASIAN
GREBE. Marshes, ponds, lakes, quiet streams. Lowlands in Java,
Timor, Sangi I. in the Sangihe Is., Ternate I. in the Moluccas; New Guinea
on the s coast and locally on n coast; Rennell I. in the s Solomon Is.;
Vanuatu on Santa Maria, Espíritu Santo and Aoba; New Caledonia;
Australia, Tasmania and New Zealand. Sympatric with ruficollis in
New Guinea, possibly in Java and the Lesser Sunda Is.
Tachybaptus rufolavatus ALAOTRA GREBE.
Lakes, marshes. Lake Alaotra in ec Madagascar. Population estimated
as ca. 20 pairs. Confined to Lake Alaotra where it hybridizes with,
and is being swamped by, T. ruficollis, a recent colonizer; rufolavatus
may be a race of ruficollis.
Tachybaptus pelzelnii MADAGASCAR GREBE.
Lakes, marshes. Lowlands to 1800 m of Madagascar.
Tachybaptus dominicus LEAST GREBE.
Marshes, lakes, ponds. Lowlands and mts. to 2700 m from s Baja Calif.,
Sinaloa and ec,s Texas s (incl. Cozumel I.) to Panama, and from Colombia,
Venezuela, Curaçao and Bonaire is., Trinidad, Tobago and Guianas
s, w of the Andes to nw Peru, and e of the Andes to e Peru, n,e,se Bolivia,
Paraguay, Uruguay and n Argentina s to Entre Ríos; Bahamas (exc.
Grand Bahama) and Greater Antilles e to the Virgin Islands.
Podilymbus podiceps PIED-BILLED GREBE.
Marshes, ponds, lakes. Lowlands and mts. to 4100 m from s coastal
Alaska in the Copper R. region (one record), and from c British Columbia,
sc Mackenzie, n Alberta, n Saskatchewan, n Manitoba, c Ontario, sw Quebec,
c Maine, s New Brunswick, Prince Edward I. and Nova Scotia s through U.S.,
Mexico, C. America and W. Indies s to Tobago and Panama, and from Colombia,
Venezuela, Trinidad and Guianas s (mostly absent from Amazonia) to c Chile
on Isla Chiloé, and s Argentina in Chubut. Recently on Hawaii.
See P. gigas.
*Podilymbus gigas ATITLAN GREBE. Probably
extinct, not recorded since 1984. Formerly only on Lake Atitlán,
c Guatemala. P. podiceps has replaced gigas on Lake Atitlán.
Poliocephalus poliocephalus HOARY-HEADED
GREBE. Swamps, lakes, estuaries, lagoons. Australia, Tasmania
and s New Zealand on the South Island. Ranges to Snares Islands.
Poliocephalus rufopectus NEW ZEALAND GREBE.
Swamps, lakes, estuaries, lagoons. New Zealand. Now reduced
in numbers and confined to North Island.
Podiceps major GREAT GREBE. Lakes,
rivers, estuaries. Coastal Peru from Piura s to Ica, Chile n to Coquimbo,
Argentina e of the Andes n to Santiago del Estero, Santa Fe and Entre Ríos,
Paraguay, Uruguay and extreme s Brazil in Rio Grande do Sul.
Podiceps grisegena RED-NECKED GREBE.
Reedy lakes, large ponds, quiet rivers. From Sweden and s Finland
se across nw,c Russia n to Ural Mts. and Ob R. drainage and se to Caspian
and Aral sea areas and n Kirghiz steppes, to w China in nw Sinkiang, s
to the Balkans, Turkey, nw Iran, Caucasus, Transcaucasus; c,e Siberia to
Kamchatka and Sakhalin, Manchuria and n Japan on Hokkaido. N. America
from w,c Alaska, c Yukon, nw,s Mackenzie, nw Saskatchewan, c Manitoba and
w,sc Ontario s to St. Lawrence I. (formerly ?), Alaska Peninsula, c Washington,
sw Oregon, n Montana, nw Wyoming, ne S. Dakota, sc Minnesota, n Michigan,
s Quebec and n New England.
Podiceps cristatus GREAT CRESTED GREBE.
Reed-bordered lakes, sluggish rivers. From British Isles and s Scandinavia
e across c,se Russia and sw,se Siberia to Mongolia, w,n China, probably
s Korea and Japan, s to nw,ne Africa from Morocco to Tunisia; Nile delta;
n Mediterranean region on Corsica and Sardinia, to Turkey, Iran, Transcaucasus,
Transcaspia, Turkestan and n India in Kashmir and Ladakh. E,s Africa
from highlands of Ethiopia, Zaire, s through rift valley lakes of nw Rwanda,
w Uganda and c Kenya to n Tanzania, occurring elsewhere in Angola, Namibia,
Zambia, Botswana, s Mozambique and S. Africa. In sw,e,se Australia
from extreme sw W. Australia and ne Queensland s to Victoria and w to se
S. Australia, Tasmania; S. Island of New Zealand. Range expanded
since 1950.
Podiceps auritus HORNED GREBE. Marshes,
ponds, lakes, streams. From Iceland, n British Isles and Scandinavia
e across c Russia and c Siberia to Kamchatka, s to Kirghiz steppes, w China
in w Sinkiang and s Siberia from Altai to Amurland, Sakhalin. N.
America from c Alaska, n Yukon, nw,s Mackenzie, s Keewatin and n Manitoba
s to e Washington, n Utah, nw Nebraska, ne Iowa, n Illinois, n Indiana,
s Ontario, s Quebec and s New England. Has recently disappeared from
most of U.S. breeding range.
Podiceps nigricollis BLACK-NECKED GREBE.
Large reedy lakes, ponds, saltmarsh, lagoons. From British Isles,
s Sweden, n Norway and continental Europe e across c Russia to w,ne China
and sw,se Siberia; s to n Tunisia, Spain, c France, Balkans, Turkey, Iraq,
e Iran, Trancaucasus and Kazakhstan. E,s Africa in c Ethiopia, Uganda
and rift valley lakes of c Kenya and n Tanzania; w Namibia, se Botswana,
w Zimbabwe and S. Africa. From sc British Columbia, s Yukon, c Alberta,
c Saskatchewan, sw Manitoba, w Minnesota and ne Illinois s to n Baja Calif.,
c Arizona, c,ne New Mexico, sc Texas and c Oklahoma, and Mexico s to Jalisco
and Puebla.
*Podiceps andinus COLOMBIAN GREBE. Probably extinct. Formerly on lakes and ponds in the Andes, 2500-3100 m of ec Colombia. Often included in nigricollis.
Podiceps occipitalis SILVERY GREBE. Marshy ponds, lakes, rarely bays.
The two subspecies are sometimes treated as separate
species.
P. o. juninensis. Andes, 2500-5000
m from sw Colombia s through Ecuador, Peru and c,sw Bolivia to n Chile
s to Antofagasta and nw Argentina in Jujuy and Salta. Possibly a
separate species.
P. o. occipitalis. Lowlands and mts.
to 2800 m from nc Chile n to Atacama and w,s Argentina n to San Juan, Córdoba
and Santa Fe, s to Tierra del Fuego; Falkland Islands.
Podiceps taczanowskii PUNA GREBE.
Lakes, ponds. High Andes, 4050-4100 m on Lake Junín in c Peru.
Closely related to occipitalis, but they are sympatric on Lake Junín.
Population ca. 50 birds in 1993, due to habitat degradation of Lake Junin
(Valqui, T. 1994. Cotinga, vol. 1).
Podiceps gallardoi HOODED GREBE.
Lakes in windswept uplands. S Argentina, known only from several
lakes in c Santa Cruz Province.
Aechmophorus occidentalis WESTERN GREBE.
Lakes, bays, rarely saltmarsh. From sc British Columbia, c Alberta,
c Saskatchewan and sw Manitoba s to s Calif., nc Utah, sw Colorado, sw,ne
New Mexico, w Nebraska, nw Iowa and w Minnesota and ec Wisconsin; n Mexico
from Chihuahua and Durango s to n Guerrero, Puebla and San Luis Potosí.
The two "color morphs" -- light phase and dark phase -- in populations of Aechmophorus are now considered to be two species, A. occidentalis and A. clarkii. They are widely sympatric, but clarkii is rare in the north and dominant in the south. DNA hybridization data indicate a difference comparable to congeneric species.
Aechmophorus clarkii CLARK'S GREBE. Lakes, bays, rarely saltmarsh. W North America, rare is parts of same breeding range as the Western Grebe, s to Nayarit and Guanajuato.
Order CICONIIFORMES
Suborder Ciconii
Infraorder Ciconiides
Parvorder Phaethontida
*Family
Phaethontidae: Tropicbirds
Parvorder Sulida
Superfamily
Suloidea
*Family
Sulidae: Boobies, Gannets
*Family
Anhingidae: Anhingas
Superfamily
Phalacrocoracoidea
*Family
Phalacrocoracidae: Cormorants
Parvorder Ciconiida
Superfamily
Ardeoidea; Family Ardeidae: Herons, Egrets, Bitterns
Superfamily
Scopoidea; Family Scopidae: Hamerkop or Hammerhead Stork
Superfamily
Phoenicopteroidea; Family Phoenicopteridae: Flamingos
Superfamily
Threskiornithidae; Family Threskiornithidae: Ibises, Spoonbills
Superfamily
Pelecanoidea
Family Pelecanidae
Subfamily Balaenicipitinae: Shoebill
*Subfamily Pelecaninae: Pelicans
Superfamily
Ciconioidea: New World Vultures, Storks
Superfamily
Procellarioidea
*Family
Fregatidae: Frigatebirds
Family Spheniscidae: Penguins
Family Gaviidae: Loons or Divers
Family Procellariidae: Petrels, Albatrosses, Shearwaters, Fulmars, etc.
The groups preceded by an asterisk (*) are members of the traditional Order Pelecaniformes
Order CICONIIFORMES:
Family PHAETHONTIDAE:
Phaethon aethereus RED-BILLED TROPICBIRD.
Pelagic, breeding on islands on cliff ledges or seaside caves. Islands
in Gulf of California and on Consag Rock and San Pedro Mártir and
San Jorge islands; Revillagigedo, Tres Marías and Isabela islands
and Alijos Rocks off w Mexico, and islands off S. America incl. Galapagos
Is. and islands from Colombia s to c Peru. Caribbean on Culebra and
Vieques is. off Puerto Rico; small islets in the Virgin Is. and Lesser
Antilles s to Grenada and Tobago; on Swan Key in Almirante Bay, nw Panama;
and off n Venezuela on Los Hermanos and Los Roques. Islands off Africa
incl. Cape Verde Is., Ascension, St. Helena and Socotra, and small islands
off Senegal, Eritrea, Djibouti, Somalia and Yemen. Islands off Brazil;
islands in the Red Sea, Gulf of Aden and Persian Gulf. Ranges at
sea in the Atlantic n to New York and s to Brazil; in w Pacific to the
Paracel Islands in the s China Sea; e Pacific n to Washington, w to Hawaiian
Is., and s to Chile; in tropical Indian Ocean.
Phaethon rubricauda RED-TAILED TROPICBIRD. Pelagic, breeds on islands on cliffs, in craters and on sand. Tropical islands from the Bonin and Volcano islands off s Japan, e to w Hawaiian Is. e to Kauai, Lanai, Kahoolawe; Manana I. off Oahu; s to Raine I. off ne Australia, and from Lord Howe, Norfolk and Kermadec is., e to Easter I. and probably Sala-y-Gómez. Indian Ocean on islands off sw Madagascar, near Mauritius, in Aldabra, Seychelles and Cocos Is. on Keeling I.; Gunungapi in the Lesser Sundas, and on islands off nw Australia. Ranges in Pacific n to Japan and s to New Zealand; in Indian Ocean n to Red Sea and Persian Gulf and s to S. African and Australian waters.
Phaethon lepturus WHITE-TAILED TROPICBIRD. Pelagic, breeds on islands on cliff ledges, caverns and trees. Islands in the Pacific from Bonin and Volcano is. off s Japan e to Hawaiian Is. on the main islands w to Kauai, rarely on Midway, s to New Caledonia, Fiji, Tonga on Eau I., Marquesas and Tuamotu. Atlantic-Caribbean region from Bermuda and W. Indies s to islets off Tobago, Fernando de Noronha off Brazil, Ascension I., and islands in Gulf of Guinea. Indian Ocean from Seychelles and Andaman Is. s to Mascarenes and Christmas I. Ranges at sea through tropical oceans n to Japan and Nova Scotia, and s to Australia, New Zealand, n S. America and S. Africa.
Family SULIDAE:
Papasula abbotti ABBOTT'S BOOBY.
Pelagic. Breeds in trees on islands. Christmas I. in the e
Indian Ocean; formerly Assumption I., Mascarenes on Mauritius and Rodrigues,
and Chagos Arch. on Gloriosa I. Ranges at sea in the Indian Ocean.
Subfossils from the Solomon Is. and Marquesas Is. represent populations
that may have survived past 1700 (Schubel and Pahlavan. 1988. Proc. Biol.
Soc. Wash. 101:487).
Morus bassanus NORTHERN GANNET. Pelagic.
Breeds on open islands and coastal cliffs. Islands in the n Atlantic
in Gulf of St. Lawrence on Bonaventure, Anticosti and Magdalen is., formerly
off Quebec; Newfoundland, formerly Nova Scotia; and New Brunswick; islands
around Iceland, Faroes, British Isles, n France and coastal Norway.
Ranges at sea in n Atlantic s to Gulf coast, Cape Verde Is., nw Africa
and Mediterranean Sea, and nw to n Russia and the Baltic Sea.
Morus capensis CAPE GANNET. Pelagic.
Breeds on cliffs. Islands off s Africa from Hollandsbird I. off Namibia,
s and e to e S. Africa at Algoa Bay. Ranges at sea n to Gulf of Guinea
and Kenya. May be a subspecies of bassanus.
Morus serrator AUSTRALIAN GANNET.
Pelagic. Breeds on open ground on islands. Islands off se Australia
in s Victoria incl. Lawrence Rock and Wedge Light; off ne,s Tasmania on
Black Pyramid, Cat I., Pedra Blanca, Eddystone Rock; New Zealand on islets
off n N. Island, at Cape Kidnappers on se North I., and off se S. Island.
Ranges at sea along coasts of s Australia n to c W. Australia and se Queensland,
and in New Zealand waters. May be a race of bassanus.
Sula nebouxii BLUE-FOOTED BOOBY. Pelagic.
Breeds on open ground on islands. Islands off Pacific coast in Gulf
of California n to Consag Rock and Tres Marietas Is., off Pacific Honduras
on Los Farollones, on Pearl Is. and small islets in Gulf of Panama; Galapagos
Islands; islands along coast from Colombia to Peru. Ranges at sea
in e Pacific from Baja Calif. s to s Peru.
Sula variegata PERUVIAN BOOBY. Pelagic.
Breeds on cliffs. Islands off Pacific coast from nw Peru s to c Chile
to Concepción. Ranges at sea from sw Colombia to s Chile and
north to Bay of Panama in El Niño years.
Sula dactylatra MASKED BOOBY. Pelagic.
Breeds on open ground on oceanic islands. Is. off Mexico on Alijos
Rocks, Revillagigedo, Clipperton; through tropical Pacific from Ryukyu
and Hawaiian is. on Kure e to Kaula, and on Moku Manu off Oahu; s to is.
in the Sulu Sea (Tubbataha Reef), off se Queensland; Kermadec Is., Tuamotu
and Sala-y-Gómez; Galapagos and islands from Malpelo I. off Colombia
s to San Ambrosio and San Félix off Chile. Atlantic-Caribbean
region on islands off Yucatán Peninsula; Florida Keys on the Dry
Tortugas; s Bahamas, sw of Jamaica, off Puerto Rico, Virgin Is. and Lesser
Antilles off Anguilla and in the Grenadines; Isla Los Monjes off n Colombia
and Islas Aves e to Los Hermanos off Venezuela; from off Brazil to Ascension
I. Is. off w Africa in Gulf of Guinea, off Cameroon. Indian
Ocean from Red Sea, Gulf of Aden, coast of Eritrea, Somalia, Socotra Is.,
se Arabia off s Oman, Keeling I. in the Cocos Is., and Christmas Is. s
to islands off Kenya, Tanzania, Zanzibar, Seychelles, Comoro and Mascarene
is. Islands off nw Australia and Gunungapi I. in the Lesser Sundas.
Ranges widely in tropical oceans, rarely n to Gulf coast of se U.S. and
N. Carolina, and s to w Mexico, se Australia and S. Africa.
Sula sula RED-FOOTED BOOBY. Pelagic. Breeds in bushes and trees on islands. Islands in Pacific on the Paracel Is. in the s China Sea; from Bonin Is. and Hawaiian Is. from Kure e to Kauai, Oahu and Moku Manu, s to Tubbataha Reef in the Sulu Sea, and to ne Australia on Raine I., New Caledonia, Fiji, Samoa, Tonga on Kalau I., Tuamotu and Sala-y-Gómez; Revillagigedo and Tres Marías off Mexico; Cocos I. off Costa Rica; Isla La Plata off Ecuador; Galapagos Is. Atlantic-Caribbean region on Glover's Reef off Yucatán; Half Moon Cay off Belize; Swan Is. off Puerto Rico; Virgin Is. and Grenadines, off Tobago and islands off Venezuela from Los Roques e to Los Hermanos and off Brazil on Fernando de Noronha and Trindade. Indian Ocean from Socotra Is., Aldabra Is., Cosmoledo, Europa, Farquhar and Tromelin is. off Madagascar, Comoros, Seychelles, e to Keeling in Cocos Is., and Gunungapi in the Lesser Sundas. Ranges at sea from Sinaloa to Panama, from Gulf coast (rarely) s along Middle and S. America to e Brazil and in Indian Ocean n to Bay of Bengal.
Sula leucogaster BROWN BOOBY. Pelagic. Breeds on open ground and cliffs on islands. Pacific islands from Izu, Bonin, Volcano and Hawaiian is., from Kure e to Kauai and Moku Manu, s to islands in S. China Sea; Tubbataha Reef in the Sulu Sea; off n Australia, New Caledonia, Tonga and Tuamotu; islands in Gulf of California; off w Mexico on the Revillagigedo, Clipperton, Tres Marías and Isabela is., probably on Los Farallones off Honduras; islets off Costa Rica; Pearl Is. and islets in Bay of Panama and Gorgona I. off Colombia. Atlantic-Caribbean region from islands off Yucatán Pen., formerly Florida Keys; and Bahamas s through Antilles and along coast of e Middle and n S. America e to Trinidad; from Cape Verde Is., islands off w Africa and Gulf of Guinea s to coast of c Brazil and Ascension I.; islands in the Indian Ocean from Red Sea, Gulf of Aden and Malay Peninsula, s to Seychelles, Mitsio I. off Madagascar; Keeling I., and Christmas Is.; and Gunungapi in the Lesser Sundas. Ranges at sea n, rarely,to Japan, Calif. and Nova Scotia, s, rarely, to Ecuador, S. Africa and se Australia.
FamilyANHINGIDAE:
Anhinga anhinga ANHINGA. Swamps,
lakes, sluggish streams, lagoons, mangroves. Lowlands to 500 m from
c,e Texas, se Oklahoma, s,e Arkansas, s Missouri (formerly), s Illinois
(formerly), w Tennessee, c Gulf states and coastal N. Carolina s to Florida;
from Sinaloa and s Texas s through lowlands on both slopes to Panama, and
from Colombia, Venezuela, Trinidad, Tobago and Guianas s, w of the Andes
to w Ecuador and e of Andes through e Ecuador, e Peru, n,e Bolivia, Brazil
and Paraguay to Uruguay and n Argentina; locally in Cuba, Isle of Pines
and Grenada in the s Lesser Antilles.
Anhinga rufa AFRICAN DARTER. Swamps,
lakes, sluggish streams, marshes. From sw Mauritania and Senegambia
e through s Mali, Guinea-Bissau, Guinea, Sierra Leone to Ghana, s Niger,
Nigeria, s Chad and c,s Sudan to Ethiopia and Somalia, s to Gabon and Zaire
and to Rwanda, Burundi, Uganda, Kenya, Tanzania, Angola, Zambia, Malawi,
Mozambique, Namibia, Botswana and Zimbabwe to s S. Africa; Madagascar.
Breeds in most African countries and occurs in virtually all in suitable
habitats, possibly breeding in others in addition to those listed above.
Anhinga melanogaster ORIENTAL DARTER. Swamps, marshes, lakes, flooded fields. Lowlands to 300 m in s Turkey (formerly), se Iraq, and from India, Sri Lanka e to Burma, Thailand, Indochina and Malaya (vagrant?); Sumatra, Java, Borneo, Sulawesi and Philippines on Luzon, Mindoro, Palawan, Negros and Mindanao.
Relationships among the Old World darters are uncertain. A. melanogaster is closest to novaehollandiae and they are often considered conspecific and sometimes not even subspecifically distinct. A. rufa is next closest and the Old World populations may constitute a single species. A. anhinga is the most distant from the others, but the entire genus may be a single species. They are treated here as allospecies.
Anhinga novaehollandiae AUSTRALIAN DARTER. Rivers, lakes, swamps, lagoons, estuaries. Lowlands in New Guinea, Fergusson I. in the D'Entrecasteaux Arch., and locally in Australia. Ranges to the Lesser Sunda Is. and s Moluccas.
Family PHALACROCORACIDAE:
*
Siegel-Causey (1988. Condor 90:885-905) revised
the Phalacrocoracidae and placed the following species in two subfamilies:
Phalacrocoracine to include the "Cormorants" from P. africanus through
P. capillatus and Leucocarboninae to include the "Shags" from P. nigrogularis
to P. featherstoni.
The 37 species were divided among nine genera.
DNA hybridization values approximately support the clustering of species
proposed by Siegel-Causey.
The recognition of nine genera seems inflated
relative to other groups, but this is a matter of opinion.
*
Phalacrocorax africanus LONG-TAILED CORMORANT.
Rivers, lakes, swamps. Coasts and inland waters from sw Mauritania,
Senegambia, s Mali, s Niger, São Tomé I., s Chad, c,s Sudan,
Ethiopia and w,s Somalia s to s S. Africa; Madagascar. Occurs and
breeds in most African countries with suitable habitat. Not recorded
as breeding in Djibouti, Socotra Is., Yemen, Seychelles, Aldabra, Comores
or Mascarene Is. Wanders widely to offshore islands.
Phalacrocorax coronatus CROWNED CORMORANT.
Rocky islands, seacoasts. From c Angola in Benguela s to Namibia
and sw S. Africa to Cape Agulhas. May be conspecific with africanus.
Phalacrocorax pygmeus PYGMY CORMORANT.
Marshes, reedbeds, lakes, rivers, seacoasts. From Balkans n to Yugoslavia
and Romania, Greece and w Turkey e to shores of Black, Caspian and Aral
seas, n,sw Tadshikistan, se Iraq and n Iran.
Phalacrocorax niger LITTLE CORMORANT.
Lakes, rivers, marshes, mangroves. Lowlands from India, Sri Lanka
and sw China s through se Asia (exc. Malaya); n Java.
Phalacrocorax melanoleucos LITTLE PIED
CORMORANT. Coasts, islands, reefs, estuaries, lakes, marshes, ponds,
swamps, etc. Lowlands and mts. to 3400 m in the East Indies (exc.
Sumatra and Borneo) incl. many small islands; w Papuan Is. of Waigeo, Salawati
and Misool; New Guinea incl. Numfor I., Palau, Solomon and Santa Cruz islands;
New Caledonia; Australia, Tasmania. New Zealand, Stewart I., and
Campbell I. P. m. brevirostris (New Zealand) is sometimes treated
as a separate species.
*Phalacrocorax perspicillatus PALLAS'S
CORMORANT. Extinct by ca. 1850. Known from five specimens from
Bering I. in the Commander Is.
Phalacrocorax penicillatus BRANDT'S CORMORANT.
Coasts, bays. S coastal Alaska in Prince William Sound; from Washington
s to Baja Calif. in I. Natividad, San Cristobal Bay, formerly on Guadelupe
I.; islands in Gulf of California on San Pedro Mártir, Salsipuedes,
Roca Blanca.
Phalacrocorax harrisi FLIGHTLESS CORMORANT.
Coasts of the Galapagos islands of Fernandina and Isabela (exc. s,sw).
Phalacrocorax neglectus BANK CORMORANT.
Coasts, weedy banks. Coastal Namibia n to Walvis Bay and sw S. Africa
in sw Cape Province.
Phalacrocorax fuscescens BLACK-FACED CORMORANT.
Coasts, islands, bays, rocky headlands. From sw W. Australia w to
Arch. Recherche and Hopetoun, e to extreme se New S. Wales; Tasmania and
most islands in Bass Strait.
Phalacrocorax brasilianus NEOTROPIC CORMORANT.
Rivers, lakes, swamps, coasts. Lowlands and foothills to 2600 m from
s Sonora, New Mexico, nc,e Texas and sw Louisiana s (incl. is. off Yucatán
Pen.) to Panama, and from Colombia, Venezuela, Trinidad and Guianas s to
Tierra del Fuego. Reported breeding in Cuba, Isle of Pines and Bahamas
is questionable. Formerly called P. olivaceus, but brasilianus has
priority.
Phalacrocorax auritus DOUBLE-CRESTED CORMORANT.
Lakes, swamps, rivers, coasts. From s,se Alaska w to Cape Peirce
and e Aleutians, coastal British Columbia, n Alberta and from c Saskatchewan
e across s Canada to Gulf of St. Lawrence and Newfoundland, s to Baja Calif.,
coastal Sonora, sw Arizona, s New Mexico, nc,se Texas, Gulf coast, Florida,
n Bahamas s to Cat I., San Salvador and Great Inagua; Cuba and Isle of
Pines; formerly on Man-of-War Key off Belize.
Phalacrocorax fuscicollis INDIAN CORMORANT.
Marshes, lakes, rivers, swamps. Lowlands in Pakistan in Sind, India
n to Himalayan foothills, Sri Lanka, s Burma, c,se Thailand, Cambodia and
s Vietnam in Cochinchina.
Phalacrocorax varius PIED CORMORANT.
Estuaries, bays, coasts, lakes, rivers. Locally in Australia and
New Zealand incl. Stewart I.
Phalacrocorax sulcirostris LITTLE BLACK
CORMORANT. Coasts, bays, mangroves, rivers, swamps, lakes.
Lowlands to 1200 m in the E. Indies (exc. Sumatra, but incl. many small
islands), New Guinea, Fergusson I., Australia, Tasmania and North I. of
New Zealand.
Phalacrocorax carbo GREAT CORMORANT.
Lakes, rivers, coasts, marshes.
P. c. carbo. From s Greenland, Iceland,
Faroes, British Isles, Norway and extreme nw Russia on the Kola Pen., s
to France e to Poland; s to n Africa from Mauritania e to Egypt, Sardinia,
Corsica, w coast of Italy, Czechoslovakia, Balkans, Greece and Turkey e
to Black and Caspian seas, s Russia, Mongolia, China and s Siberia to Korea
and Japan on Honshu, s to n Iran, Pakistan, India to 3500 m, Sri Lanka
and se Asia (exc. Malaya) (formerly Sumatra); n Philippines on Batan Is.,
Calayan, Luzon and Ticao. Australia, Tasmania, New Zealand and Chatham
Is. Ne N. America along Atlantic coast from n shore of Gulf of St.
Lawrence and sw Newfoundland s to Prince Edward I., Nova Scotia and n Maine.
P. c. lucidus. Senegambia, sw Mauritania,
Burkina Faso, Guinea-Bissau, Ivory Coast, Ghana, Nigeria, Cameroon, L.
Chad area, and from Gabon and Congo to s Angola, Namibia, Zambia, ne,ce,se
Zaire, Sudan, Ethiopia and s Somalia s to Rwanda, Burundi, Uganda, Kenya,
Tanzania, Malawi, Zimbabwe, Botswana, Mozambique and s S. Africa.
Phalacrocorax capillatus JAPANESE CORMORANT.
Rocky coasts and islands. Se Siberia in Ussuriland, s Kuril Is.,
Korea, ne China and Japan s to n,e Hokkaido and n Honshu.
Phalacrocorax nigrogularis SOCOTRA CORMORANT.
Coasts, islands. Islands in the Persian Gulf, se coast of Arabia
and Socotra I. Ranges to Gulf of Aden and Red Sea.
Phalacrocorax capensis CAPE CORMORANT.
Coasts, bays. Namibia n to Walvis Bay and s S. Africa in s Cape Province,
possibly off Angola, Gabon and Mozambique.
Phalacrocorax bougainvillii GUANAY CORMORANT.
Coasts, islands. Coastal islets from n Peru n to Piura s to c Chile
at Santiago (formerly Mocha I.); se Argentina in Chubut.
Phalacrocorax verrucosus KERGUELEN SHAG.
Islands, coastal cliffs. Kerguelen Is., s Indian Ocean.
Phalacrocorax atriceps IMPERIAL SHAG.
Coasts, cliffs, lakes. S Chile n to Isla Mocha and s Argentina n
to Chubut, and on lakes in Neuquén and Río Negro; Falkland
Is. In s Indian Ocean on Heard, Marion, Crozet, Kerguelen and Macquarie
islands.
"P. albiventer" appears to be a morph of P. atriceps. P. georgianus and P. bransfieldensis are often considered races of P. atriceps, although bransfieldensis may have been or still is sympatric with atriceps in extreme s S. America (Siegel-Causey and Lefevre 1989. Condor 91:408-415). P. verrucosus and the P. carunculatus-campbelli complex are similar and may also belong to this assemblage.
Phalacrocorax bransfieldensis ANTARCTIC
SHAG. Coasts, cliffs, lakes. Subantarctic islands of S. Orkney
and S. Shetland; Graham Land on the Antarctic Peninsula.
Phalacrocorax georgianus SOUTH GEORGIA
SHAG. Coasts, cliffs, lakes. S. Georgia and S. Sandwich islands.
Phalacrocorax campbelli CAMPBELL ISLAND
SHAG. Rocky islets, cliffs. Campbell I. May be conspecific
with carunculatus. The systematics of the large shags of the New
Zealand region remains unclear. The several forms are variously included
in one species or divided into six species, as here, following Siegel-Causey
(1988. Condor 90:885-905), who reported minor skeletal differences based
on small samples. P. campbelli may even be a race of P. atriceps.
Phalacrocorax carunculatus ROUGH-FACED SHAG.
Rocky islets. Islands in Cook Strait, c New Zealand.
Phalacrocorax chalconotus BRONZE SHAG.
Rocky islets. Islands on coastal se South Island of New Zealand in
Otago, and Stewart Island.
Phalacrocorax onslowi CHATHAM ISLANDS SHAG.
Rocky islets. Chatham Is.
Phalacrocorax colensoi AUCKLAND ISLANDS
SHAG. Rocky islets, cliffs. Auckland Is.
Phalacrocorax ranfurlyi BOUNTY ISLANDS
SHAG. Rocky islets, cliffs. Bounty Is.
Phalacrocorax magellanicus ROCK SHAG.
Coasts, rocky shores. From c Chile n to Valdivia and se Argentina
n to Chubut, s to Cape Horn; Falkland Is.
Phalacrocorax urile RED-FACED CORMORANT.
Coasts, rocky islands, coastal cliffs. Islets off e Hokkaido in n
Japan; Commander, Pribilof and Aleutian islands, and along coast of s Alaska
on Cape Peirce, Walrus I. and Prince William Sound.
Phalacrocorax kenyoni AMCHITKA CORMORANT.
Described from abundant recent and subfossil skeletal material from Amchitka
I., Alaska, and from three skeletonized specimens collected in 1958.
The status of kenyoni will be in doubt until it can be identified in life
and compared with P. urile, which it closely resembles, and with P. pelagicus.
See Auk 110:680, 1993.
Phalacrocorax pelagicus PELAGIC CORMORANT.
Coasts, cliffs. E China s to Kwantung, Korea, Japan off Hokkaido
and n Honshu; e Siberia from Wrangel I. and Chukotski Pen., s to Ussuriland
and Sakhalin, incl. Kamchatka and Commander Is.; from Alaska n to Cape
Lisburne and Aleutians, and British Columbia s to n Baja Calif. on Los
Coronados Is.
Phalacrocorax aristotelis EUROPEAN SHAG.
Coasts. From w Iceland, British Isles, Norway and the Kola Pen. in
nw Russia, s along coasts of w Europe to Mediterranean and Black seas,
incl. most islands and the African coast, and Atlantic coast of Morocco.
The African race, riggenbachi, sometimes is treated as a species.
Phalacrocorax gaimardi RED-LEGGED CORMORANT.
Coasts, cliffs. From Peru n to Piura, s to s Chile in the Straits
of Magellan, and coastal s Argentina in Santa Cruz.
Phalacrocorax punctatus SPOTTED SHAG.
Coasts, cliffs, rocky islands. New Zealand on the N. Island, w S.
island and Stewart I.
Phalacrocorax featherstoni PITT ISLAND
SHAG. Coasts, cliffs, rocky islands. Chatham Islands.
Often treated as a race of P. punctatus.